Catalogue of the type fossils stored in the Palaeontological Museum of "La Sapienza" University of Rome: 2

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INTRODUCTION

This is the second contribution towards an inventory of all the type fossils stored in the Palaeontological Museum of “La Sapienza” University of Rome. The types are all refigured here. For each type the main data are given and subsequent designations are not reported.

COLLECTIONS

Three are the main type-bearing collections here considered: AGIP, Almera and Cerio collections.

The AGIP collection is constituted by about eighty echinoid specimens, found in Bathonian, Cenomanian and Eocene sediments of Somalia and Ethiopia. This material was studied by Checchia-Rispoli (1943, 1945, 1950) and Maccagno (1941, 1946, 1947). AGIP is the acronym of “Azienda Generale Italiana Petroli” (=

General Italian Company of Petroleum). The researchers of this company, directed by prof. C. Migliorini, worked in the African colonies of Italy between the years 1936- 1938 for petroleum research.

The Almera collection stored in the Rome paleonto- logical Museum is constituted by forty specimens of Cretaceous corals found in Catalonia (Spain). This material was studied by De Angelis d’Ossat (1905b). Jaume Almera (1845-1919) was priest, professor of geology and natural history in the Seminario Conciliar de Barcelona. In the year 1874 he founded the “Museo de Geognosia y Paleontología del Seminario de Barcelona”.

Besides he became president of the “Real Academia de Ciencias y Artes de Barcelona” in 1907.

The Cerio collection is constituted by a dozen of Cretaceous coral specimens found near Venassino, in the Capri island. These corals were studied by De Angelis d’Ossat (1905a). Ignazio Cerio (1841-1921) was an eclectic personage: physician, scientist and archaeo- logist. He lived at Capri for fifty years, collecting shells, fossils, minerals, plants and animals.

THE CATALOGUE ORGANISATION

This catalogue is taxonomically organised: within each mayor grouping, the taxa are listed by alphabetic order.

For each type there are eight records:

Record 1 - genus and species name, followed by the name of author(s) and year of institution of the species;

Record 2 - inventory number of MPUR (Museo Pale- ontologia Università Roma). The inventory numbers are prefixed i. (invertebrate), ve. (vertebrate) and vg. (plant);

Record 3 - name of the collection;

Record 4 - kind of type: holotype or syntype or lectotype;

Record 5 - stratigraphic distribution

Record 6 - geographic distribution (country, region, locality). For Italian localities the initials of the province name are put between brackets, after the type locality name;

Record 7 - original bibliographic reference, with the number of the illustration of the type specimen;

Record 8 - remarks.

The types

All the specimens utilized for describing a new species are named types. In particular the holotype is the specimen, chosen by the author, to represent a new species. The remaining specimens of that new species are identified as paratypes. The type nomenclature is regulated by the ICZN (International Code of Zoological Nomenclature) (1999).

If the holotype was not designed, all the specimens examined to describe a new species are syntypes. Among the syntypes, a later author may choose a lectotype. In this case the remaining syntypes are named paralecto- types.

MUSEUM OF “LA SAPIENZA” UNIVERSITY OF ROME: 2 Riccardo Manni

Dipartimento di Scienze della Terra, Università degli Studi di Roma “La Sapienza”, P.le A. Moro 5, I-00185 Roma, Italy riccardo.manni@uniroma1.it

ABSTRACT - This paper is a second contribution towards a catalogue of all types stored in the Palaeontological Museum of “La Sapienza” University of Rome. The present list deals mainly with the echinoids and corals.

KEY WORDS: holotypes, syntypes, catalogue of types, Palaeontological Museum, Anthozoa, Echinoidea, Crinoidea.

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Amphiastraea waltheri De Angelis d’Ossat, 1905 Fig. 1

MPUR: i. 407 Coll. Cerio holotype

Cretaceous, Barremian

Italy, Campania, Venassino (NA)

De Angelis d’Ossat (1905a): Pl. 1, Figs. 9, 10.

Axosmilia almerai De Angelis d’Ossat, 1905 Fig. 2

MPUR: i. 443 Coll. Almera syntype

Cretaceous, Aptian

Spain, Catalonia, C. Pascual

De Angelis d’Ossat (1905b): Pl. 16, Figs. 5c-d.

Axosmilia bofilli De Angelis d’Ossat, 1905 Fig. 3

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 16, Fig. 4a.

Chaetetes capri 1 De Angelis d’Ossat, 1905 Fig. 4

MPUR: i. 404 Coll. Cerio syntype

Cretaceous, Barremian

Italy, Campania, Venassino (NA)

De Angelis d’Ossat (1905a): Pl. 1, Figs. 1, 2.

Remarks: this subspecies was identified by De Angelis (1905a) with number 1. This specimen has been polished.

INVERTEBRATA

COELENTERATA ANTHOZOA

Fig. 1 - Amphiastraea waltheri De Angelis d’Ossat, 1905 Holotype MPUR: i. 407 (x 0.3).

Fig. 2 - Axosmilia almerai De Angelis d’Ossat, 1905 Syntype MPUR: i. 443 (x 1.6).

Fig. 3 - Axosmilia bofilli De Angelis d’Ossat, 1905 Syntype MPUR: i. 434 (x 1).

Fig. 4 - Chaetetes capri 1 De Angelis d'Ossat, 1905 Syntype MPUR: i. 404 (x 0.9).

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Peplosmilia casañasi De Angelis d’Ossat, 1905 Fig. 5

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 17, Figs. 5c-d.

Peplosmilia coquandi De Angelis d’Ossat, 1905 Fig. 6

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 17, Fig. 2b.

Remarks: the specimen figured in Pl. 17, Fig. 7 (De Angelis d’Ossat, 1905b) has a small cut on the left side.

Peplosmilia fromenteli De Angelis d’Ossat, 1905 Fig. 7

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 17, Figs. 6b-c.

Remarks: This specimen has been polished.

Peplosmilia thildae De Angelis d’Ossat, 1905 Fig. 8

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Morgades De Angelis d’Ossat (1905b): unfigured.

Remarks: only this unfigured specimen of this species is stored in this Museum. This specimen has been polished.

Fig. 5 - Peplosmilia casañasi De Angelis d'Ossat, 1905 Syntype MPUR: i. 439 (x 1).

Fig. 6 - Peplosmilia coquandi De Angelis d'Ossat, 1905 Syntype MPUR: i. 436 (x 0.9).

Fig. 7 - Peplosmilia fromenteli De Angelis d'Ossat, 1905 Syntype MPUR: i. 440 (x 1).

Fig. 8 - Peplosmilia thildae De Angelis d'Ossat, 1905 Syntype MPUR: i. 435 (x 1.2).

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Peplosmilia iberica De Angelis d’Ossat, 1905 Fig. 9

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 17, Fig. 4c.

Thecosmilia catalaunica De Angelis d’Ossat, 1905 Fig. 10

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 14, Fig. 6b.

Trochosmilia n. sp. 3 De Angelis d’Ossat, 1905 Fig. 11

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 15, Fig. 6a.

Remarks: this species was identified by De Angelis d’Ossat (1905b) without specific name but only as “n.

sp. 3”.

Trochosmilia nevianii De Angelis d’Ossat, 1905 Fig. 12

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 15, Fig. 3d.

Fig. 9 - Peplosmilia iberica De Angelis d'Ossat, 1905 Syntype MPUR: i. 438 (x 1).

Fig. 10 - Thecosmilia catalaunica De Angelis d'Ossat, 1905 Syntype MPUR: i. 416 (x 0.65).

Fig. 11 - Trochosmilia n. sp. 3 De Angelis d'Ossat, 1905 Syntype MPUR: i. 431 (x 1.2).

Fig. 12 - Trochosmilia nevianii De Angelis d'Ossat, 1905 Syntype MPUR: i. 425 (x 1.7).

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Trochosmilia portisi De Angelis d’Ossat, 1905 Fig. 13

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 15, Fig. 2f.

Trochosmilia sandalina De Angelis d’Ossat, 1905 Fig. 14

Cretaceous, Aptian

Spain, Catalonia, Castellví de la Marca, C. Pascual De Angelis d’Ossat (1905b): Pl. 15, Fig. 4d.

Brissoides dainellii Checchia-Rispoli, 1943 Fig. 15

MPUR: i. 477 Coll. AGIP syntype

Paleogene, Eocene

Somalia, Migiurtina (= Bari Region)

Checchia-Rispoli (1943): Pl. 1, Fig. 5, Fig. 7.

Remarks: this specimen was figured by Checchia-Rispoli (1943) in Pl. 1, Fig. 5 and Fig. 7 as mirror images.

Diplopodia gigantea Checchia-Rispoli, 1945 Fig. 16

MPUR: i. 162 Coll. AGIP holotype

Cretaceous, Cenomanian Somalia, Uebi Scebeli

Checchia-Rispoli (1945): Pl. 1, Fig. 5.

Remarks: Checchia-Rispoli (1945) described this species in the note 2 of pag. 88.

Fig. 13 - Trochosmilia portisi De Angelis d'Ossat, 1905 Syntype MPUR: i. 423 (x 1.9).

Fig. 14 - Trochosmilia sandalina De Angelis d'Ossat, 1905 Syntype MPUR: i. 427 (x 1.1).

Fig. 15 - Brissoides dainellii Checchia-Rispoli, 1943 Syntype MPUR: i. 477 (x 1).

Fig. 16 - Diplopodia gigantea Checchia-Rispoli, 1945 Holotype MPUR: i. 162 (x 0.6).

ECHINODERMATA ECHINOIDEA

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Dorocidaris cerullii Checchia-Rispoli, 1923 Fig. 17

MPUR: i. 13

Coll. Checchia-Rispoli holotype

Neogene, Pliocene

Italy, Latium, coast between Anzio and Nettuno (Rome) Checchia-Rispoli (1923): Pl. 2, Fig. 5.

Remarks: this specimen has lost some plates.

Echinopygus checchiai Maccagno, 1947 Fig. 18

Jurassic, Bathonian

Somalia, Migiurtina (= Bari Region), Almedò Maccagno (1947a): Pl. 1, Fig. 14, Figs. 14a-b.

Remarks: this specimen was figured by Maccagno (1947a) in Pl. 1, Fig. 14 as mirror image.

Echinus portisi Checchia-Rispoli, 1923 Fig. 19

MPUR: i. 8

Neogene, Pliocene

Italy, Latium, coast between Anzio and Nettuno (Rome) Checchia-Rispoli (1923): Pl. 1, Fig. 1.

Gymnocidaris gortanii Maccagno, 1946 Fig. 20

MPUR: i. 614 Coll. AGIP - Gortani holotype

Jurassic, Dogger.

Ethiopia, Harar, Mount Darima Assa Maccagno (1946): Pl. 2, Fig. 15, Figs. 15a-b.

Remarks: the specimen corresponds to that indicated by Maccagno (1946) as “diameter 20 mm, heigth 1.10 mm and mouth diameter 0.95 mm”.

Fig. 17 - Dorocidaris cerullii Checchia-Rispoli, 1923 Holotype MPUR: i. 13 (x 0.7).

Fig. 18 - Echinopygus checchiai Maccagno, 1947 Holotype MPUR: i. 462 (x 0.7).

Fig. 19 - Echinus portisi Checchia-Rispoli, 1923 Holotype MPUR: i. 8 (x 0.35).

Fig. 20 - Gymnocidaris gortanii Maccagno, 1946 Holotype MPUR: i. 614 (x 1.75).

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Hemiaster gemellaroi Checchia-Rispoli, 1936 Fig. 21

MPUR: i. 156 Coll. Trevisan holotype

Cretaceous, Cenomanian Italy, Sicily, Caltavuturo (PA)

Checchia-Rispoli (1936): Pl. 16, Fig. 5, Fig. 5a.

Hemidiadema mortenseni Maccagno, 1947 Fig. 22

MPUR: i. 396 Coll. AGIP syntype

Jurassic, Bathonian?

Somalia, Migiurtina (= Bari Region), Bogol Magmo?

Maccagno (1947a): Pl. 1, Fig. 6b.

Remarks: the specimen was figured by Maccagno (1947a) in Pl. 1, Fig. 6 as mirror image. The age and the locality are doubtful.

Kleinia pulcra Checchia-Rispoli, 1950 Fig. 23

Paleogene, Eocene

Somalia, Migiurtina (= Bari Region), Carin Checchia-Rispoli (1950): Pl. 2, Fig. 4, Fig. 4a.

Remarks: this specimen was figured by Checchia- Rispoli in Pl. 2, Fig. 4 as mirror image.

Linthia mortenseni depressa Checchia-Rispoli, 1950 Fig. 24

Paleogene, Eocene

Somalia, Migiurtina (= Bari Region), Galgalò Checchia-Rispoli (1950): Pl. 1, Fig. 5, Figs. 5a-b.

Remarks: this specimen was figured by Checchia- Rispoli (1950) in Pl. 1, Fig. 5b as mirror image.

Fig. 21 - Hemiaster gemellaroi Checchia-Rispoli, 1936 Holotype MPUR: i. 156 (x 1.65).

Fig. 22 - Hemidiadema mortenseni Maccagno, 1947 Syntype MPUR: i. 396 (x 2.4).

Fig. 23 - Kleinia pulcra Checchia-Rispoli, 1950 Holotype MPUR: i. 472 (x 1.4).

Fig. 24 - Linthia mortenseni depressa Checchia-Rispoli, 1950 Holotype MPUR: i. 476 (x 1.4).

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Periaster maugerii Checchia-Rispoli, 1936 Fig. 25

MPUR: i. 155 Coll. Trevisan holotype

Cretaceous, Cenomanian Italy, Sicily, Caltavuturo (PA)

Pliolampas trevisani Checchia-Rispoli, 1936 Fig. 26

MPUR: i. 153 Coll. Trevisan syntype

Paleogene, Eocene Italy, Sicily, Pachino (SR)

Prospatangus distefanoi praecursor Checchia-Rispoli, 1923

Fig. 27 MPUR: i. 18

Coll. Checchia-Rispoli syntype

Neogene, Pliocene

Italy, Latium, coast between Anzio and Nettuno (Rome) Checchia-Rispoli (1923): Pl. 4, Fig. 1, Fig. 1a.

Psammechinus macrostomus Checchia-Rispoli, 1932 Fig. 28

MPUR: i. 132

Coll. Checchia-Rispoli syntype

Neogene, Pliocene

Italy, Latium, Anzio (Rome)

Checchia-Rispoli (1932): Pl. 1, Figs. 3, 4.

Fig. 25 - Periaster maugerii Checchia-Rispoli, 1936 Holotype MPUR: i. 155 (x 1.15).

Fig. 26 - Pliolampas trevisani Checchia-Rispoli, 1936 Syntype MPUR: i. 153 (x 1.75).

Fig. 27 - Prospatangus distefanoi praecursor Checchia-Rispoli, 1923 Syntype MPUR: i. 18 (x 0.3).

Fig. 28 - Psammechinus macrostomus Checchia-Rispoli, 1932 Syntype MPUR: i. 132 (x 1.65).

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Pseudocidaris gortanii Maccagno, 1947 Fig. 29

Jurassic, Bathonian

Somalia, Migiurtina (= Bari Region), Comina Maccagno (1947a): Pl. 1, Fig. 8.

Remarks: this specimen was figured by Maccagno (1947a) in Pl. 1, Fig. 9 as mirror image.

Pseudocidaris migliorinii Maccagno, 1947 Fig. 30

Jurassic, Bathonian

Somalia, Migiurtina (= Bari Region), Bogol Magmo Maccagno (1947a): Fig. 2; Pl. 1, Fig. 7, Fig. 7a.

Remarks: this specimen was figured Maccagno (1947a) by in Fig. 2 and in Pl. 1, Fig. 7, Fig. 7a as mirror images.

Schizaster aichinoi Checchia-Rispoli, 1923 Fig. 31

MPUR: i. 20

Neogene, Pliocene

Italy, Latium, coast between Anzio and Nettuno (Rome) Checchia-Rispoli (1923): Pl. 3, Fig. 2, Fig. 2a.

Schizaster alsiensis Maccagno, 1947 Fig. 32

MPUR: i.134

Neogene, Upper Pliocene Italy, Latium, Palo (Rome)

Maccagno (1947b): Pl. 1, Figs. 1-4.

Fig. 29 - Pseudocidaris gortanii Maccagno, 1947 Holotype MPUR: i. 452 (x 2.65).

Fig. 30 - Pseudocidaris migliorinii Maccagno, 1947 Holotype MPUR: i. 399 (x 1.9).

Fig. 31 - Schizaster aichinoi Checchia-Rispoli, 1923 Holotype MPUR: i. 20 (x 0.5).

Fig. 32 - Schizaster alsiensis Maccagno, 1947 Holotype MPUR: i.134 (x 0.75).

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Fig. 33 MPUR: i. 473

Coll. AGIP syntype

Paleogene, Eocene

Somalia, Migiurtina (= Bari Region), rigth side of the Uadi

Checchia-Rispoli (1950): Pl. 2, Fig. 5, Figs. 5a-c.

Remarks: the specimen corresponds to that indicated by Checchia-Rispoli (1950) as “length 26 mm, width 22.3 and heigth 19.5 mm”; the specimen was figured by Checchia-Rispoli (1950) in Pl. 2, Fig. 5a and 5c as mirror images.

Somaliaster magniventer checchiai Maccagno, 1941 Fig. 34

Paleogene, Eocene

Somalia, Migiurtina (= Bari Region), Hardàg Maccagno (1941): Pl. 11, Figs. 10, 11.

Phyllocrinus checchiai Serra, 1933 Fig. 35

MPUR: i. 160 Coll. Maugeri holotype

Cretaceous, Neocomian

Italy, Sicily, Licodia Eubea, Boschitello Mounts (SR) Serra (1933): Figs. 1-3.

Fig. 33 - Schizaster delorenzoi Checchia-Rispoli, 1950 Syntype MPUR: i. 473 (x 1.4).

Fig. 34 - Somaliaster magniventer checchiai Maccagno, 1941 Holotype MPUR: i. 493 (x 1.1).

Fig. 35 - Phyllocrinus checchiai Serra, 1933 Holotype MPUR: i. 160 (x 4.1).

CRINOIDEA

REFERENCES

Checchia-Rispoli G. (1923) - Gli echinidi del Pliocene di Anzio. Mem. R. Com. Geol. It., 9, 6-29.

Checchia-Rispoli G. (1932) - Sopra uno Psammechinus del Pliocene di Anzio (Roma). Boll. R. Uff. Geol. It., 57 (12), 1-6.

Checchia-Rispoli G. (1936) - Su alcuni echinidi della Sicilia.

Boll. Soc. Geol. It., 55 (2), 295-310.

Checchia-Rispoli G. (1943) - Brissoidi eocenici della Migiurtina. Mem. R. Accad. Naz. Lincei, s. 6, 14, 99-111.

Checchia-Rispoli G. (1945) - Di due nuovi generi di echinidi del Cretaceo della Somalia. Boll. R. Uff. Geol. It., 68 (8) [1943], 81-90.

Checchia-Rispoli G. (1950) - Su alcuni echinidi eocenici della Migiurtina. Boll. Uff. Geol. It., 70 [1945-1946], 21-41.

De Angelis d’Ossat G. (1905a) - I coralli del calcare di Venassino (isola di Capri). Atti R. Acc. Sc. Fis. Mat.

Napoli, s. 2, 12 (16), 1-45.

De Angelis d’Ossat G. (1905b) - Coralli del Cretacico inferio- re della Catalogna. Palaeontographia Italica, 11, 169-252.

International Commission on Zoological Nomenclature (1999) - International Code of Zoological Nomenclature adopted by the International Union of Biological Sciences. Fourth edition. 306 pp. London.

Maccagno A.M. (1941) - Osservazioni sul genere Somaliaster

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Maccagno A.M. (1946) - Echinidi giurassici dell’Harar. Boll.

R. Uff. Geol. It., 69 (4) [1944], 79-101.

Maccagno A.M. (1947a) - Illustrazione degli echinidi giurassi- ci della Somalia. Mem. Accad. Naz. Lincei, s. 8, 1, 99-136.

Maccagno A.M. (1947b) - Nuova specie di Schizaster nel Macco di Palo (Lazio). Boll. Soc. Geol. It., 65 [1946], 115- 121.

Neocomiano della Sicilia. Boll. Soc. Geol. It., 52 (2), 353- 356.

Accettato per la stampa: Settembre 2007

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