Scuola Dottorale di Ateneo
Graduate School
Dottorato di ricerca
in Scienze Ambientali
Ciclo XXIX
Anno di discussione 2017
Trophic niche variability of fish populations and
communities as a response to ontogeny, habitat
heterogeneity and restoration
SETTORE SCIENTIFICO DISCIPLINARE DI AFFERENZA
: BIO/07
Tesi di Dottorato di Cristina Andolina, matricola 956081
Coordinatore del Dottorato
Tutore del Dottorando
Prof. Gabriele Capodaglio
Prof. Piero Franzoi
Co-tutore del Dottorando
Prof.ssa Salvatrice Vizzini
“When we try to pick out anything by itself,
we find it hitched to everything else in the universe”
1
NŸŘǼsŘǼǣ
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2
ÝŘǼNjŸ_ȖOǼÞŸŘ
ȕŘ_sNjǣǼŘ_ÞضŘǼȖNjĶɚNjÞEÞĶÞǼɴŸ¯sOŸĶŸ¶ÞOĶƼNjŸOsǣǣsǣÞǣONjȖOÞ͝ŸNjÞŘǼsNjƼNjsǼÞض sOŸǣɴǣǼsŎ¯ȖŘOǼÞŸŘÞضŘ_OÌÞsɚÞض¯ÞNjŎضsŎsŘǼƼNjOǼÞOsǣʹĵŘ_NjsǣʰōŸNj¶Ř ˓ ǢɠŘǣŸŘ ˠ˨˨˨ʺʳ ōŸNjsŸɚsNjʰ ÞǼ ĶĶŸɠǣ ǼŸ ÞŘǼsNjƼNjsǼ NjsǣƼŸŘǣs ǼŸ _ÞǣǼȖNjEŘOsʰ _ÞǣOsNjŘÞضǼÌsEǣsĶÞŘsŸ¯ŘǼȖNjĶɚNjÞEÞĶÞǼɴ¯NjŸŎĶǼsNjs_ƼNjŸOsǣǣsǣʹ®NjǼsNjNjÞ¶Ÿ˓ NJȖǣĨˡ˟˟˧ʱɳŸȖضsǼĶʳˡ˟ˠˤʺʳ ŗǼȖNjĶɚNjÞEÞĶÞǼɴÞŘsŘɚÞNjŸŘŎsŘǼĶǣɴǣǼsŎǣNjÞǣsǣ¯NjŸŎÞŘǼNjÞŘǣÞO_ɴŘŎÞOǣŸ¯EŸǼÌ EÞŸǼÞOŘ_EÞŸǼÞOƼNjŸOsǣǣsǣʹ®NjǼsNjNjÞ¶Ÿ˓NJȖǣĨˡ˟˟˧ʺʳƻNjÞŎNjÞĶɴǼÌsɚNjÞEÞĶÞǼɴŸ¯ EÞŸǼÞO¯OǼŸNjǣʰOŸŘǣÞ_sNjs_sÞǼÌsNjǼǼÌsǣƼǼÞĶǣOĶsʹsʳ¶ʳ¶Nj_ÞsŘǼǣʰÌsǼsNjŸ¶sŘsÞǼɴʺ ŸNjǼsŎƼŸNjĶŸŘsʹsʳ¶ʳǣsǣŸŘĶÞǼɴʺÞŘ_ȖOsǣŸNj¶ŘÞǣŎǣǼŸ_ƼǼǼŸŘsɠOŸŘ_ÞǼÞŸŘǣʰ ɠÌÞOÌÞŘ¯ĶȖsŘOsĶĶǼÌsǣƼsOǼǣŸ¯ǼÌsĶÞ¯sOɴOĶsŸ¯ŘŸNj¶ŘÞǣŎŘ_ÞǼǣEsÌɚÞŸȖNjʰ ¯NjŸŎ¶NjŸɠǼÌʹNJȖǣǣŸʰNŸǣǼ˓NǼȖ_sĶĶˡ˟˟˦ʺǼŸNjsƼNjŸ_ȖOǼÞŸŘʹNɚNjNjŸʰəNjÞŘ˓ ōĶɚǣÞˡ˟ˠˣʺʰÞŘOĶȖ_Þض¯ss_ÞضÌEÞǼǣŘ_NjsǣŸȖNjOsȖǣsʹsʳ¶ʳ®NjŘOÞǣsǼĶʳʰˡ˟ˠˠʺʰ ÌEÞǼǼOÌŸÞOsŘ_ÞŘǼsNjOǼÞŸŘǣɠÞǼÌŸǼÌsNjǣƼsOÞsǣʹsʳ¶ʳ^ŘǼǣsǼĶʳˡ˟ˠˡʱōȖŘĨsǼ Ķʳˡ˟ˠˣʺʳǢsOŸŘ_ĶɴʰEÞŸǼÞOƼNjŸOsǣǣsǣǣȖOÌǣÞŘǼsNj˚Ř_ÞŘǼNj˚ǣƼsOÞ¯ÞOOŸŎƼsǼÞǼÞŸŘʰ _ȖsǼŸ¯ŸŸ_Ř_ǣƼOs ĶÞŎÞǼǼÞŸŘʰŎÞŘĶɴNjs¶ȖĶǼsǼÌsEȖŘ_ŘOsŸ¯ƼŸƼȖĶǼÞŸŘǣ ʹsʳ¶ʳNNjssǣs˓ȕŘ_sNjɠŸŸ_ˠ˨˧ˡʺʳǻÌsÞŘǼsNjƼĶɴŸ¯ǣȖOÌ_ɴŘŎÞOǣʰǼŸ¶sǼÌsNjɠÞǼÌǼÌs ÞŘǼNjÞŘǣÞOĶÞ¯sÌÞǣǼŸNjɴǼNjÞǼǣʰOŸŘǼNjÞEȖǼsǣǼŸǣǼNjȖOǼȖNjsǼÌsɚNjÞEÞĶÞǼɴǼ_Þ¯¯sNjsŘǼĶsɚsĶǣ Ÿ¯ ŸNj¶ŘÞʊǼÞŸŘʰ ¯NjŸŎ ÞŘ_ÞɚÞ_ȖĶǣ ǼŸ ƼŸƼȖĶǼÞŸŘǣ ǼŸ OŸŎŎȖŘÞǼÞsǣ ʹƻŘǼsĶsǼ Ķʳ ˡ˟ˠˣʺʳ ŘɴŸNj_sNjŸ¯ƼsNjǼȖNjEǼÞŸŘʰsÞǼÌsNjŘs¶ǼÞɚsʹÞŎƼOǼʺŸNjƼŸǣÞǼÞɚsʹsʳ¶ʳʰŎضsŎsŘǼ ŸNj NjsǣǼŸNjǼÞŸŘ ƼNjOǼÞOsǣʺʰ ÞŘ_ȖOsǣ ǣŸŎs OÌضs ÞŘ ǼÌs _ɴŘŎÞOǣ ŎsŘǼÞŸŘs_ʰ NjsǣȖĶǼÞض ÞŘ ǣǼNjȖOǼȖNjĶ ɚNjÞǼÞŸŘ Ÿ¯ ƼŸƼȖĶǼÞŸŘǣ Ř_ OŸŎŎȖŘÞǼÞsǣ ÞŘɚŸĶɚs_ʰ ɠÌÞOÌ OŘNjsŎÞŘNjsĶǼÞɚsĶɴǣǼEĶsɠÞǼÌÞŘOsNjǼÞŘǼŸĶsNjŘOsNjضsǣʰŸNjOŘsɚsŘ_ɚŘOsŸNj Njs¶NjsǣǣǼŸŸŘsŸ¯ǼÌsĨŘŸɠŘǣǼǼsǣŸ¯ŘsOŸĶŸ¶ÞOĶǣȖOOsǣǣÞŸŘʹ®NjǼsNjNjÞ¶Ÿ˓NJȖǣĨ ˡ˟˟˧ʺʳ3 ÝŘǼÌsĶǣǼ_sO_sǣʰÞŘONjsǣÞضǼǼsŘǼÞŸŘÌǣEssŘƼŸǣs_ǼŸǼNjŸƼÌÞOÞŘǼsNjOǼÞŸŘǣǣ Ȗǣs¯ȖĶ ǼŸŸĶǣ ǼŸ ÞŘɚsǣǼÞ¶Ǽs ǼÌs _s¶Njss Ÿ¯ OŸŎƼĶsɮÞǼɴ Ř_ ǣǼEÞĶÞǼɴ Ÿ¯ ǣɴǣǼsŎ ʹDsͶNjŘŸsǼĶʳ ˡ˟˟ˤʱ^ȖŘŘs ˡ˟˟˥ʺʰ ǣ ǼÌsɴNjsƼNjsǣsŘǼ ŸŘs Ÿ¯ ǼÌsĨsɴ ÞŘǼsNjOǼÞŸŘ ǼɴƼŸĶŸ¶ÞsǣǼÌǼOŘŸOOȖNjEsǼɠssŘǣƼsOÞsǣʹħw¯ÞsǼĶʳˡ˟ˠˡʺʳ ǢÞŘOssOŸĶŸ¶ÞǣǼǣÌɚsĶɠɴǣǼNjÞs_ǼŸÞ_sŘǼÞ¯ɴǼÌsNjŸĶsŸ¯ǣƼsOÞsǣÞŘǼÌsOŸŘǼsɮǼŸ¯ ǼÌsÞNj OŸŎŎȖŘÞǼɴʰ Eɴ ǣȖŎŎNjÞʊÞض ĶĶ ǼÌs ǣƼsOǼǣ Ÿ¯ ǼÌsÞNj ĶÞ¯s ÌÞǣǼŸNjɴ ǼNjÞǼǣ Ř_ ÌEÞǼǣ ʹNŸÌsŘ ˠ˨˦˦ʺʰ ǼÌs OŸŘOsƼǼ Ÿ¯ ǼNjŸƼÌÞO ŘÞOÌs Ìǣ EsOŸŎs ŎÞĶsǣǼŸŘs ÞŘ ǼNjŸƼÌÞO ǣǼNjȖOǼȖNjs ǣǼȖ_Þsǣ ʹrĶǼŸŘ ˠ˨ˡ˦ʺʳ ǻÌs ǼNjŸƼÌÞO ŘÞOÌs NjsƼNjsǣsŘǼǣ ŸŘs Ÿ¯ ǼÌs _Þ¯¯sNjsŘǼ ǣƼsOǼǣ Ÿ¯ ǼÌs ŎŸNjs OŸŎƼNjsÌsŘǣÞɚs OŸŘOsƼǼ Ÿ¯ sOŸĶŸ¶ÞOĶ ŘÞOÌsʰ ɠÌÞOÌ ÞŘOĶȖ_sǣǼÌsŸɚsNjĶĶNjsLJȖÞNjsŎsŘǼǣŘss_s_¯ŸNjǣƼsOÞsǣǼŸƼsNjǣÞǣǼŘ_NjsƼNjŸ_ȖOsÞŘ ¶ÞɚsŘsŘɚÞNjŸŘŎsŘǼʳǻÌÞǣOŘEsÞŘǼsNjƼNjsǼs_ÞŘǼsNjŎǣŸ¯ǣƼOsˀÌEÞǼǼŸOOȖƼÞs_ ʹµNjÞŘsĶĶ˅ǣ ŘÞOÌsʺʰ ¯ŸŸ_ ÌEÞǼǣ ʹrĶǼŸŘ˅ǣ ŘÞOÌsʺ Ř_ OŘ Es ÞŎ¶ÞŘs_ ǣ Ř Ř˚ _ÞŎsŘǣÞŸŘĶ ÌɴƼsNjɚŸĶȖŎs ɠÌsNjs ɮsǣ Njs ¶ÞɚsŘ Eɴ sŘɚÞNjŸŘŎsŘǼĶ ɚNjÞEĶsǣ ʹËȖǼOÌÞŘǣŸŘ˂ǣŘÞOÌsʺʳ
ƻNjǼÞOȖĶNjĶɴʰǼÌsǼNjŸƼÌÞOŘÞOÌsɠǣŸNjÞ¶ÞŘĶĶɴ_s¯ÞŘs_EɴrĶǼŸŘʹˠ˨ˡ˦ʺǣǼÌsǣȖŎŸ¯ ĶĶǼÌsǼNjŸƼÌÞOÞŘǼsNjOǼÞŸŘǣŸOOȖNjNjÞضŎŸŘ¶ǣƼsOÞsǣÞŘǣɴǣǼsŎʳŗÞOÌsƼNjŎsǼsNjǣ Njs ÞŘ¯ĶȖsŘOs_ Eɴ Řɴ ÞŘǼNj˚ Ř_ ÞŘǼsNjǣƼsOÞ¯ÞO OÌضs Ÿ¯ ǼNjŸƼÌÞO ÞŘǼsNjOǼÞŸŘǣ ʹDsNjÌŸƼ sǼ Ķʳ ˡ˟˟ˣʺʰ ÌsŘOs ǼÌs ŘĶɴǣÞǣ Ÿ¯ ǼNjŸƼÌÞO ǣǼNjȖOǼȖNjs OŘ ƼNjŸɚÞ_s LJȖŘǼÞǼǼÞɚsĶɴ ÞŘ¯ŸNjŎǼÞŸŘ ŸŘ ǼÌs ǼNjŸƼÌÞO ɚNjÞEÞĶÞǼɴ Ÿ¯ ǣƼsOÞsǣʰ ɠÌÞOÌ OŘ Es ǣǣsǣǣs_sÞǼÌsNjǼƼŸƼȖĶǼÞŸŘŸNjOŸŎŎȖŘÞǼɴĶsɚsĶǣʳ ĵŸŸĨÞضǼǼÌsƼŸƼȖĶǼÞŸŘĶsɚsĶʰŎȖOÌŸ¯ǼÌsɚNjÞEÞĶÞǼɴÞŘ¯ss_ÞضÌEÞǼǣʰNjsǣŸȖNjOs ȖǣsŘ_ÌEÞǼǼOÌŸÞOsNjsĶÞŘĨs_ǼŸǼÌsǼNjŸƼÌÞO_ƼǼEÞĶÞǼɴŸ¯ǼÌsǣƼsOÞsǣʰɠÌÞOÌʰ OOŸNj_ÞضǼŸµsNjĨÞضʹˠ˨˨ˣʺʰ_s¯ÞŘsǣǼÌsEÞĶÞǼɴŸ¯ŘŸNj¶ŘÞǣŎǼŸEsŘs¯ÞǼ¯NjŸŎǼÌs ŎŸǣǼ¯ɚŸȖNjEĶs¯ŸŸ_ǣŸȖNjOsɚÞĶEĶsÞŘƼNjǼÞOȖĶNjsŘɚÞNjŸŘŎsŘǼʰǼƼNjǼÞOȖĶNj ǼÞŎsʳǻÌs¶NjsǼsNjǼÌsEÞĶÞǼɴ¯ŸNjǣƼsOÞsǣǼŸǣÌÞ¯ǼǼŸǼÌsȖǣsŸ¯_Þ¯¯sNjsŘǼǣŸȖNjOsǣʰ ǼÌs ¶NjsǼsNj ǼÌs ƼŸǣǣÞEÞĶÞǼɴ ǼŸ ƼsNjǣÞǣǼ ÞŘ ¶ÞɚsŘ sŘɚÞNjŸŘŎsŘǼ ȖŘ_sNj OÌضÞض OŸŘ_ÞǼÞŸŘǣʰEɴƼNjǼÞǼÞŸŘÞضǼÌsǣŸȖNjOsǣɚÞĶEĶsÞŘOǣsŸ¯OŸŎƼsǼÞǼÞŸŘɠÞǼÌŸǼÌsNj ǣƼsOÞsǣʹDNjNjŸǣsǼĶʳˡ˟ˠˢʺʳ
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ǻNjŸƼÌÞO_ƼǼEÞĶÞǼɴÞǣǼɴƼÞO͝ŸNjsɮŎƼĶsŸ¯ŸƼƼŸNjǼȖŘÞǣǼÞO¯ÞǣÌǣƼsOÞsǣÞŘÌEÞǼÞض sOŸǣɴǣǼsŎǣOÌNjOǼsNjÞʊs_EɴÌÞ¶ÌŘǼȖNj͝ĶȖOǼȖǼÞŸŘǣŸ¯sŘɚÞNjŸŘŎsŘǼĶɚNjÞEĶsǣʰ ǣȖOÌǣǼNjŘǣÞǼÞŸŘĶsOŸǣɴǣǼsŎǣʹəÞʊʊÞŘÞ˓ōʊʊŸĶˡ˟˟ˡʱəÞضNjssǼĶʳˡ˟ˠˡʱħŸƼƼ sǼĶʳˡ˟ˠˢʺʳ®ŸNjǼÌÞǣNjsǣŸŘʰǣȖO̯ÞǣÌǣƼsOÞsǣOŘEsOŸŘǣÞ_sNjs_ǣŎŸ_sĶǣǼŸǣǼȖ_ɴ ǼÌs _s¶Njss Ÿ¯ ɚNjÞEÞĶÞǼɴ Ÿ¯ ǼÌs ǼNjŸƼÌÞO ŘÞOÌs ɠÞǼÌÞŘ ǼÌs ǣŎs ǣɴǣǼsŎʰ EsǼɠssŘ _Þ¯¯sNjsŘǼǣɴǣǼsŎǣŸNjONjŸǣǣǼÞŎsʳ
rǣǼȖNjÞsǣʰOŸǣǼĶͶŸŸŘǣŘ_ƼŸŘ_ǣNjsǼNjŘǣÞǼÞŸŘĶsOŸǣɴǣǼsŎǣʰsOŸǼŸŘsǣĶŸOǼs_ Ǽ ǼÌs ÞŘǼsNj¯Os EsǼɠssŘ ǼÌs ŎNjÞŘs Ř_ ǼÌs ǼsNjNjsǣǼNjÞĶ _ŸŎÞŘǣʳ ^Ȗs ǼŸ ǼÌsÞNj ƼŸǣÞǼÞŸŘʰ ǼÌs ŎÞɮÞض Ÿ¯ OŸŘǼÞŘsŘǼĶ ɠǼsNjǣ ɠÞǼÌ ŎNjÞŘs ɠǼsNjǣ _sǼsNjŎÞŘsǣ ǼÌs ÞŘǣǼEÞĶÞǼɴ Ÿ¯ ƼÌɴǣÞOŸ˛OÌsŎÞOĶ ɚNjÞEĶsǣ ǣȖOÌ ǣ ǣĶÞŘÞǼɴʰ ǼsŎƼsNjǼȖNjsʰ Ÿɮɴ¶sŘʰ ɠÌŸǣs¯ĶȖOǼȖǼÞŸŘsɮǼsŘǼ_sƼsŘ_ǣĶǣŸŸŘǼÞ_sʰsɚƼŸNjǼÞŸŘʰNjÞŘ¯ĶĶǣŘ_¯ĶŸŸ_Þضǣʰ Ř_ NjsǣȖĶǼǣ ÞŘ ÌÞ¶Ì sŘɚÞNjŸŘŎsŘǼĶ ÌsǼsNjŸ¶sŘsÞǼɴ ʹNŸ¶ŘsǼǼÞʰ ǢNj ˓ ō¶ʊʊȘ ˡ˟˟˧ʱ ƻwNjsʊ˚NJȖʊ¯sǼ Ķʳ ˡ˟ˠ˟ʺʳ ŷŘ ǼÌs ŸǼÌsNj ǣÞ_sʰ ÌŸɠsɚsNjʰ ǣȖOÌ sŘɚÞNjŸŘŎsŘǼĶ ÌsǼsNjŸ¶sŘsÞǼɴŘ_ɚNjÞEÞĶÞǼɴNjsǣȖĶǼǣÞŘÌÞ¶ÌÌEÞǼǼÌsǼsNjŸ¶sŘsÞǼɴʰǼÌǼǼŸ¶sǼÌsNj ɠÞǼÌǼÌsÌÞ¶ÌƼNjŸ_ȖOǼÞɚÞǼɴOÌNjOǼsNjÞʊÞضǼÌsǣsǣɴǣǼsŎǣǣȖƼƼŸNjǼĶNj¶s_ÞɚsNjǣÞǼɴŸ¯ EÞŸĶŸ¶ÞOĶ OŸŎŎȖŘÞǼÞsǣ ǼÌǼ ǼÌNjÞɚs Ř_ OŸŘǼNjÞEȖǼs ǼŸ ŎÞŘǼÞŘ ÌÞ¶Ì ĶsɚsĶǣ Ÿ¯ EÞŸ_ÞɚsNjǣÞǼɴʰŘŸǼŸŘĶɴɠÞǼÌÞŘǼÌsǣŎsǣɴǣǼsŎʰEȖǼĶǣŸÞŘǼÌs_ĠOsŘǼŸŘsǣʰŎNjÞŘs Ř_ sǣǼȖNjÞŘsʰ ƼĶɴÞض ONjȖOÞĶ NjŸĶs ÞŘ ǼÌs OŸŘŘsOǼÞɚÞǼɴ ŎŸŘ¶ ǣɴǣǼsŎǣ ʹʹǻ¶ĶÞƼÞsǼNjsǼ Ķʳ ˡ˟ˠˣʺʳ ®ȖNjǼÌsNjŎŸNjsʰ ǼÌs _Þ¯¯sNjsŘǼ ǼɴƼŸĶŸ¶Þsǣ Ÿ¯ ÌEÞǼǼǣ Njs Ȗǣs_ Eɴ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ ¯ŸNj ǣsɚsNjĶ NjsǣŸŘǣʰ ÞŘOĶȖ_Þض ǣƼɠŘÞضʰ ŘȖNjǣsNjɴ Ř_ ¯ss_Þض ¶NjŸȖŘ_ǣ ʹ®NjŘOŸʰ ǻŸNjNjÞOsĶĶÞ ˓ ®NjŘʊŸÞ ˡ˟˟˨ʺʳ ĶǼÌŸȖ¶Ì _sǣONjÞEÞض ǼÌs ĶÞ¯s ǣǼNjǼs¶Þsǣ Ÿ¯ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ ǼÌǼ ÞŘÌEÞǼ ǣȖOÌ OŸǣǼĶ˚ŎNjÞŘs Ř_ sǣǼȖNjÞŘs sŘɚÞNjŸŘŎsŘǼǣ ʹǣsŘǣȖ ®NjŘOŸsǼ Ķʳ ˡ˟˟˧ʺ ÌsĶƼǣ ǼŸ ȖŘ_sNjǣǼŘ_ ǼÌs Ȗǣs Ǽ ǣƼǼÞĶ Ř_ ǼsŎƼŸNjĶ ǣOĶsǣ Ÿ¯ ɚÞĶEĶs NjsǣŸȖNjOsǣʰ Njs¯ĶsOǼÞض ǼÌs sOŸǣɴǣǼsŎ ¯ȖŘOǼÞŸŘ Ÿ¯ ǼÌsǣssŘɚÞNjŸŘŎsŘǼǣʰŎŸNjsLJȖŘǼÞǼǼÞɚsǣǼȖ_ÞsǣŸ¯ǣȖOÌȖǣsNjsŘsOsǣǣNjɴʳ
ǻÌsOÌضÞضOŸŘ_ÞǼÞŸŘǣ¶ÞɚsŘEɴÌÞ¶ÌÌEÞǼǼÌsǼsNjŸ¶sŘsÞǼɴǣÌƼsǼÌsĶÞ¯sÌÞǣǼŸNjɴ ǼNjÞǼǣʹDNjÞ¶ŸĶÞŘsǼĶʳˡ˟ˠ˥ʺʰŘ_ǣOŸŘǣsLJȖsŘOsʰǼÌsɴNjsƼNjs_ÞOǼs_ǼŸǣÌƼsĶǣŸ ǼÌsǼNjŸƼÌÞOǣǼNjȖOǼȖNjsʰÌsŘOsǼÌsǼNjŸƼÌÞOŘÞOÌsŸ¯¯ÞǣÌƼŸƼȖĶǼÞŸŘǣʳ
5
ŎŸŘ¶ǼÌs_Þ¯¯sNjsŘǼǼsOÌŘÞLJȖsǣȖǣs_ÞŘǼNjŸƼÌÞOǣǼȖ_Þsǣʹsʳ¶ʳǣǼŸŎOÌOŸŘǼsŘǼʰ¯ǼǼɴ OÞ_ǣʺʰ ǼÌs ŘĶɴǣÞǣ Ÿ¯ ǣǼEĶs ÞǣŸǼŸƼsǣʰ sǣƼsOÞĶĶɴ Ÿ¯ ONjEŸŘ ʹˠˢNˀˠˡN ̠ δˠˢNʺ Ř_
ŘÞǼNjŸ¶sŘ ʹˠˤŗˀˠˣŗ = δˠˤŗʺʰ sŎsNj¶s_ ǣ ŸŘs Ÿ¯ ǼÌs ƼNjÞŎNjɴ ǼŸŸĶǣ ǼŸ _sǣONjÞEs ǼÌs
ǼNjŸƼÌÞOŘÞOÌsɠÞǼÌLJȖŘǼÞǼǼÞɚsƼƼNjŸOÌʳ
ƻNjsɚÞŸȖǣ ɠŸNjĨǣʰ ÞŘ ¯OǼʰ Ìɚs ǣÌŸɠŘ ÌŸɠ ǼÌs OŸŘOsƼǼ Ÿ¯ ǼNjŸƼÌÞO ŘÞOÌs OŘ Es ǼNjŘǣƼŸǣs_ÞŘǼŸÞǣŸǼŸƼÞOĨsɴʰEɴOŸŘǣÞ_sNjÞضǼÌsδ˛ǣƼOs¶ÞɚsŘEɴǼÌsEÞ˚ƼĶŸǼδˠˢN˚
δˠˤŗ ǣ ǼɠŸ _ÞŎsŘǣÞŸŘĶ ǣƼOs ɠÌsNjs ǼŸ NjsƼNjsǣsŘǼ ǼÌs ÞǣŸǼŸƼÞO ŘÞOÌs ǣƼOs ʹDsNjÌŸƼsǼĶʳˡ˟˟ˣʱŗsɠǣŸŎssǼĶʳˡ˟˟˦ʺʳǻÌÞǣĶĶŸɠǣȖǣǼŸȖǣsǼÌsÞǣŸǼŸƼÞOŘÞOÌs ǣ_sǣONjÞƼǼŸNjŸ¯ǼÌsǼNjŸƼÌÞOŘÞOÌsʰĶǼÌŸȖ¶ÌÞǼŘss_ǣǼŸEsĨsƼǼÞŘŎÞŘ_ǼÌǼǼÌsɴ NjsŘŸǼsɮOǼĶɴsLJȖÞɚĶsŘǼʹğOĨǣŸŘsǼĶʳˡ˟ˠˠʺʳÝŘ¯OǼʰ_sƼsŘ_ÞضŸŘǼÌsǣɴǣǼsŎ ÞŘɚŸĶɚs_ʰÞŘ¯ŸNjŎǼÞŸŘ¶ÞɚsŘ¯NjŸŎǼÌsǣŎsÞǣŸǼŸƼÞO_ǼOŸȖĶ_ƼNjŸɚÞ_sÞŘǣÞ¶ÌǼǣŸŘ ŸǼÌsNjǣƼsOǼǣŸ¯ǼÌssOŸĶŸ¶ÞOĶŘÞOÌsʰǣȖOÌǣǼÌsÌEÞǼǼȖǣsʹsʳ¶ʳĶǼÞǼȖ_sʰĶǼÞǼȖ_sʰ ǣssŗsɠǣŸŎssǼĶʳˡ˟˟˦ʺʳ ®ȖNjǼÌsNjsɮǼsŘǣÞŸŘŸ¯ǼÌsOŸŘOsƼǼÌɚsEssŘƼƼĶÞs_EɴĵɴŎŘsǼĶʳʹˡ˟˟˦ʺʰɠÌŸ ǣȖ¶¶sǣǼs_ ǣsNjÞsǣ Ÿ¯ ˆOŸŎŎȖŘÞǼɴ˚ɠÞ_s ŎsǼNjÞOǣˇ ǼÌǼ ƼNjŸɚÞ_s LJȖŘǼÞǼǼÞɚs sǣǼÞŎǼsŸ¯ǼÌsɠÞ_ǼÌŸ¯ǼÌsÞǣŸǼŸƼÞOŘÞOÌsʰǣȖOÌǣǼÌsOŸŘɚsɮÌȖĶĶsŘOŸŎƼǣǣÞض ĶĶǼÌsÞŘ_ÞɚÞ_ȖĶǣŸ¯ƼŸƼȖĶǼÞŸŘǣʰEȖǼĶǣŸǼÌsǼNjŸƼÌÞO_ÞɚsNjǣÞǼɴŘ_ǼÌsǼNjŸƼÌÞO Njs_ȖŘ_ŘOɴʰ ŸEǣsNjɚEĶs sÞǼÌsNj Ǽ OŸŎŎȖŘÞǼɴ ŸNj ƼŸƼȖĶǼÞŸŘ ĶsɚsĶʳ ōŸNjsŸɚsNjʰ ğOĨǣŸŘsǼĶʳʹˡ˟ˠˠʺĶǼsNjNjs¯ÞŘs_ǼÌsǣƼsOǼŸ¯ǼÌsɠÞ_ǼÌŸ¯ǼÌsÞǣŸǼŸƼÞOŘÞOÌsEɴ ƼƼĶɴÞض DɴsǣÞŘ ǣǼǼÞǣǼÞOǣ ǼŸ OĶOȖĶǼs ǣǼŘ_Nj_ sĶĶÞƼǣs Njs ǼÌǼ NjsǣȖĶǼ ŎŸNjs ƼNjsOÞǣs ǼÌŘ ǼÌs OŸŘɚsɮ ÌȖĶĶǣ ƼNjsɚÞŸȖǣ ŎsŘǼÞŸŘs_ʳ DŸǼÌ ƼƼĶÞOǼÞŸŘǣ ĶĶŸɠ ǼŸ OŸŎƼNjs _Þ¯¯sNjsŘǼ OŸŎŎȖŘÞǼÞsǣ ŸNj ƼŸƼȖĶǼÞŸŘǣ ONjŸǣǣ ǣƼOs ʹÌEÞǼǼǣʺʰ Ř_ ǼÌs ǣŎsŸŘsǣONjŸǣǣǼÞŎsʹĵɴŎŘsǼĶʳˡ˟˟˦ʺʳ
ǻÌs ÞŎ Ÿ¯ ǼÌÞǣ ƼNjŸĠsOǼ Þǣ ǼŸ OÌNjOǼsNjÞʊs ǼÌs ǼNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ NjsǣÞ_sŘǼ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ Ř_ OŸŎŎȖŘÞǼÞsǣ ȖŘ_sNj OÌضÞض OŸŘ_ÞǼÞŸŘǣ ǼɴƼÞOĶ Ÿ¯ OŸǣǼĶ˚ ŎNjÞŘs sOŸǣɴǣǼsŎǣʳ ǻÌNjss ŎÞŘ ¯OǼŸNjǣ ÞŘ_ȖOÞض ɚNjÞEÞĶÞǼɴ ǼŸ ǼNjŸƼÌÞO ŘÞOÌs ɠsNjs ǼĨsŘÞŘǼŸOOŸȖŘǼʲʹÞʺÌEÞǼǼÌsǼsNjŸ¶sŘÞǼɴʰʹÞÞʺŸŘǼŸ¶sŘsǼÞO_sɚsĶŸƼŎsŘǼŘ_ʹÞÞÞʺ ÌEÞǼǼNjsǣǼŸNjǼÞŸŘƼNjŸOsǣǣsǣʰŘ_ǼÌsɴɠsNjs_ÞǣOȖǣǣs_ÞŘǼÌNjssOÌƼǼsNjǣʲ
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- NÌƼǼsNj ˠ Þǣ ¯ŸOȖǣs_ ŸŘ ǼÌs ǼNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ NjsǣÞ_sŘǼ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ ONjŸǣǣ ÌEÞǼǼǣ Ǽ ÞŘǼsNj˚ Ř_ ÞŘǼNj˚ͶŸŸŘ ĶsɚsĶʰ ɠÞǼÌ ƼNjǼÞOȖĶNj ¯ŸOȖǣǼŸǼNj¶sǼǣƼsOÞsǣŸ¯ÌÞ¶ÌsOŸĶŸ¶ÞOĶɚĶȖsʰǼÌsōs_ÞǼsNjNjŘsŘǼŸŸǼÌONjƼ ƼÌŘÞȖǣ¯ǣOÞǼȖǣʱ
- NÌƼǼsNj ˡ ǣǣsǣǣsǣ ǼÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ ǼNjŸƼÌÞO ŘÞOÌs Ř_ ǼNjŸƼÌÞO ƼŸǣÞǼÞŸŘ ŸOOȖNjNjÞض _ȖNjÞض ŸŘǼŸ¶sŘɴ Ÿ¯Ǣʳ ȖNjǼ Ř_ ǼÌs NjŸĶs Ÿ¯ _Þ¯¯sNjsŘǼ ÌEÞǼǼǣ ÞŘ ǣȖƼƼŸNjǼÞض ǼÌs ǼNjŸƼÌÞO ƼǼÌɠɴ Ķs_Þض ǼŸ ǼÌs ǣƼsOÞsǣ ŸɚsNj sNjĶɴ ǣǼ¶sǣ Ÿ¯ _sɚsĶŸƼŎsŘǼɠÞǼÌÞŘOŸǣǼĶͶŸŸŘʱ - NÌƼǼsNj ˢ sɮŎÞŘsǣ ǼsŎƼŸNjĶ ɚNjÞǼÞŸŘ Ÿ¯ ǼÌs ǼNjŸƼÌÞO ŘÞOÌs Ÿ¯ ¯ÞǣÌ OŸŎŎȖŘÞǼɴǣǣŸOÞǼs_ǼŸǣÞǼsȖŘ_sNjǣs¶NjǣǣNjsǣǼŸNjǼÞŸŘʰÞŘŸNj_sNjǼŸsɚĶȖǼs ¯ȖŘOǼÞŸŘĶNjsOŸɚsNjɴŸ¯¯ÞǣÌOŸŎŎȖŘÞǼÞsǣ_ȖNjÞضÌEÞǼǼNjsǣǼŸNjǼÞŸŘƼNjŸOsǣǣsǣʳ
7
NÌƼǼsNjˠ
ǻNjŸƼÌÞOŘÞOÌsɚNjÞEÞĶÞǼɴŸ¯NjsǣÞ_sŘǼ¯ÞǣÌONjŸǣǣÌEÞǼǼǣǼ
ÞŘǼsNj˚Ř_ÞŘǼNj˚ͶŸŸŘĶsɚsĶʲ¯ŸOȖǣŸŘōs_ÞǼsNjNjŘsŘ
ĨÞĶĶÞ¯ÞǣÌ
EǣǼNjOǼNŸǣǼĶ Ķ¶ŸŸŘǣ Njs OÌNjOǼsNjÞʊs_ Eɴ ÌÞ¶Ì ÌEÞǼǼ ÌsǼsNjŸ¶sŘsÞǼɴ ɠÌsNjsʰ Ÿ¯ǼsŘʰ ŘǼȖNjĶ ÌEÞǼǼǣ OŸsɮÞǣǼ ɠÞǼÌ NjǼÞ¯ÞOÞĶ ŸŘsǣʰ ÌÞǣǼŸNjÞOĶĶɴ ǣsǼ ȖƼ ǼŸ ǣȖƼƼŸNjǼ ÌȖŎŘ OǼÞɚÞǼÞsǣʰ ¯ŸNj sɮŎƼĶs NjsĶǼs_ ǼŸ ǣĶǼ ƼNjŸ_ȖOǼÞŸŘʰ LJȖOȖĶǼȖNjs Ř_ ǼNj_ÞǼÞŸŘĶ ¯ÞǣÌsNjɴʳ ÞŎ Ÿ¯ ǼÌÞǣ ɠŸNjĨ Þǣ ǼŸ ǣǣsǣǣ ǼÌs ǼNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ NjsǣÞ_sŘǼ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ ONjŸǣǣ ŘǼȖNjĶ Ř_ NjǼÞ¯ÞOÞĶ ÌEÞǼǼǣ Ǽ ÞŘǼsNj˚ Ř_ ÞŘǼNj˚ͶŸŸŘ ĶsɚsĶʰ ɠÞǼÌ ƼNjǼÞOȖĶNj ¯ŸOȖǣ ǼŸ ǼNj¶sǼ ǣƼsOÞsǣ Ÿ¯ ÌÞ¶Ì sOŸĶŸ¶ÞOĶ ɚĶȖsʰ ǼÌs ōs_ÞǼsNjNjŘsŘǼŸŸǼÌONjƼƼÌŘÞȖǣ¯ǣOÞǼȖǣʳ ®ÞǣÌOŸŎŎȖŘÞǼÞsǣŘ_ĶĶǼÌsƼŸǣǣÞEĶsǣŸȖNjOsǣŸ¯ŸNj¶ŘÞOŎǼǼsNjɠsNjsǣŎƼĶs_ÞŘ ǣƼNjÞضŘ_ȖǼȖŎŘˡ˟ˠˣÞŘǼɠŸÌEÞǼǼǼɴƼŸĶŸ¶ÞsǣʰŸŘsNjǼÞ¯ÞOÞĶŘ_ŸŘsŘǼȖNjĶʰ ɠÞǼÌÞŘ ǼÌs ǢǼ¶ŘŸŘs _Þ ōNjǣĶ ʹǻNjƼŘÞʺ Ř_ ǼÌs ĵ¶ŸŸŘ Ÿ¯ əsŘÞOsʳ ËEÞǼǼǣ ǣŎƼĶs_ɠsNjsNjsǣƼsOǼÞɚsĶɴǣs¶NjǣǣŎs_Ÿɠ_ŸŎÞŘǼs_EɴNɴŎŸ_ŸOsŘŸ_Ÿǣ Ř_ŘsɚƼŸNjǼÞŸŘƼŸŘ_ÞŘǣĶǼɠŸNjĨǣɴǣǼsŎÞŘǼÌs¯ÞNjǣǼǣǼȖ_ɴNjsʰŘ_ǼɠŸǣĶǼ ŎNjǣÌsǣ Ř_ Ř NjǼÞ¯ÞOÞĶ _ÞǼOÌ ÞŘ ǼÌs ǣsOŸŘ_ ŸŘsʳ ǻNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ ɠǣ _sǣONjÞEs_ ȖǣÞض ONjEŸŘ Ř_ ŘÞǼNjŸ¶sŘ ǣǼEĶs ÞǣŸǼŸƼs ƼŸƼȖĶǼÞŸŘ Ř_ OŸŎŎȖŘÞǼɴ˚ ɠÞ_s ŎsǼNjÞOǣʰ ɠÌÞĶs DɴsǣÞŘ ŎÞɮÞض ŎŸ_sĶǣ ĶĶŸɠs_ ǼŸ ǼNjOĨ ǼÌs ŎÞŘ ŸNj¶ŘÞO ŎǼǼsNjƼǼÌɠɴǣǣȖƼƼŸNjǼÞضǼÌsǼNj¶sǼǣƼsOÞsǣʳ
ōÞŘ _Þ¯¯sNjsŘOsǣʰ EŸǼÌ Ǽ OŸŎŎȖŘÞǼɴ Ř_ ǼNj¶sǼ ƼŸƼȖĶǼÞŸŘ ĶsɚsĶʰ ŸOOȖNjNjs_ EsǼɠssŘ ǼÌs ǼɠŸ ǣǼȖ_ɴ Njsǣ Ř_ ɠÞǼÌÞŘ sOÌʰ EsǼɠssŘ NjǼÞ¯ÞOÞĶ Ř_ ŘǼȖNjĶ ÌEÞǼǼǣʳËEÞǼǼŘ_OŸŎŎȖŘÞǼɴǣǼNjȖOǼȖNjsNjsǣȖĶǼs_ǼŸÞŘ¯ĶȖsŘOsǼÌsǼNjŸƼÌÞOŘÞOÌs Ÿ¯ʳ¯ǣOÞǼȖǣǼÌǼÌsŘOsɚNjÞs_ÞŘ_sƼsŘ_sŘOsŸ¯ǼÌsǣƼsOÞ¯ÞOÌEÞǼǼʰEȖǼǼÌsɴ_Þ_ ŘŸǼ ÞŘ¯ĶȖsŘOs ǼÌs ŸɚsNjĶĶ sOŸĶŸ¶ÞOĶ NjŸĶs Ÿ¯ ǼÌs ǣƼsOÞsǣʳ ®ȖNjǼÌsNjŎŸNjsʰ ǼɠŸ ŎÞŘ ǼNjŸƼÌÞO ƼǼÌɠɴǣ ǣssŎs_ ǼŸ _NjÞɚs ǼÌs ¯ĶȖɮsǣ Ÿ¯ ŸNj¶ŘÞO ŎǼǼsNj Ķs_Þض ǼŸ ʳ ¯ǣOÞǼȖǣ ɠÞǼÌÞŘ ǼÌs ǼɠŸ ǼɴƼŸĶŸ¶Þsǣ Ÿ¯ ÌEÞǼǼ sɮŎÞŘs_ʲ ɠÌÞĶs ǼÌs ŎONjŸƼÌɴǼsǣ _s¯ÞŘÞǼsĶɴǣȖƼƼŸNjǼǼÌsǣƼsOÞsǣÞŘǼÌsŘǼȖNjĶÌEÞǼǼǣʰNjǼÞ¯ÞOÞĶÌEÞǼǼǣNjsŎÞŘĶɴ Eǣs_ŸŘǣs_ÞŎsŘǼNjɴŸNj¶ŘÞOŎǼǼsNjNjŸȖǼsǣʳ ŷȖNjŸȖǼOŸŎsǣĶĶŸɠs_ǼŸENjŸ_sŘĨŘŸɠĶs_¶sŸŘÌŸɠŘǼȖNjĶŘ_NjǼÞ¯ÞOÞĶǣɴǣǼsŎǣ ǣȖƼƼŸNjǼ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ ĶÞɚÞض ɠÞǼÌÞŘ OŸǣǼĶ Ķ¶ŸŸŘǣ Ř_ NjsǣȖĶǼǣ Njs _ÞǣOȖǣǣs_ ĶǣŸÞŘǼÌsĶÞ¶ÌǼŸ¯sOŸĶŸ¶ÞOĶÞŎƼĶÞOǼÞŸŘǣ¯ŸNjOŸǣǼĶŎضsŎsŘǼŸ¯ǣȖOÌNjǼÞ¯ÞOÞĶ ÌEÞǼǼǣǼÌǼOŸȖĶ_NjsƼNjsǣsŘǼÌŸǼǣƼŸǼ¯ŸNjOŸŘǣsNjɚǼÞŸŘƼNjÞŸNjÞǼÞsǣŸ¯ʳ¯ǣOÞǼȖǣʳ
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ˠʳˠʳ ÝŘǼNjŸ_ȖcǼÞŸŘ
ǻNjŘǣÞǼÞŸŘĶ sOŸǣɴǣǼsŎǣʰ ǣȖOÌ ǣ OŸǣǼĶ Ķ¶ŸŸŘǣʰ Njs ǼɴƼÞOĶĶɴ OÌNjOǼsNjÞʊs_ Eɴ ÌÞ¶Ì ǣƼǼÞŸ˚ǼsŎƼŸNjĶ ɚNjÞEÞĶÞǼɴ Ÿ¯ ǼÌs sŘɚÞNjŸŘŎsŘǼĶ ¯sǼȖNjsǣʳ ǻÌÞǣʰ ÞŘ ǼȖNjŘʰ sŘÌŘOsǣ ¶NjsǼ ÌEÞǼǼ ÌsǼsNjŸ¶sŘsÞǼɴʰ ǼÌǼ Þǣ ĶǣŸ _sƼsŘ_sŘǼ ŸŘ ÞŘǼNjÞŘǣÞO ¶sŸŎŸNjƼÌŸĶŸ¶ɴ Ÿ¯ ǼÌs ǣƼsOÞ¯ÞO ǣɴǣǼsŎʰ ǣȖOÌ ǣ ǼÌs _s¶Njss Ÿ¯ OŸŘŘsOǼÞŸŘ ǼŸ ǼÌs ŸƼsŘǣsŘ_ǼÌsÞŘƼȖǼŸ¯¯NjsǣÌɠǼsNjǣʹ®NjŘOŸʰ®NjŘʊŸÞ˓ǻŸNjNjÞOsĶĶÞˡ˟˟˧EʱəÞʊʊÞŘÞ ˡ˟˟˨ʺʳ^sƼsŘ_ÞضŸŘǼÌsͶŸŸŘʰǼÌsɚNjÞsǼɴŸ¯ÌEÞǼǼǣOŘÞŘOĶȖ_sǣĶǼŎNjǣÌsǣʰ ǣs¶NjǣǣŎs_ŸɠǣʰŎȖ_¯ĶǼǣʹsÞǼÌsNjɚs¶sǼǼs_ŸNjŘŸǼʺʰÞŘǼsNjǼÞ_ĶONjssĨǣʳōŸNjsŸɚsNjʰ ÞŘǼÌsǣssOŸǣɴǣǼsŎǣʰŘǼȖNjĶÌEÞǼǼǣŸ¯ǼsŘOŸsɮÞǣǼɠÞǼÌNjǼÞ¯ÞOÞĶŸŘsǣʰÌÞǣǼŸNjÞOĶĶɴ ǣsǼ ȖƼ ǼŸ ǣȖƼƼŸNjǼ ÌȖŎŘ OǼÞɚÞǼÞsǣ NjsĶǼs_ ¯ŸNj sɮŎƼĶs ǼŸ ǣĶǼ ƼNjŸ_ȖOǼÞŸŘʰ LJȖOȖĶǼȖNjs Ř_ NjǼÞǣŘĶ ¯ÞǣÌsNjɴʰ ɠÌÞOÌ ŎĨsǣ ǼÌsŎ ÞŎƼŸNjǼŘǼ OŸŘǼNjÞEȖǼŸNjǣ Ÿ¯ sOŸǣɴǣǼsŎǣsNjɚÞOsǣʳ
Ř sɮǼsŘǣÞɚs EŸ_ɴ Ÿ¯ NjsǣsNjOÌ Ìǣ ǣǣsǣǣs_ ǼÌs ǣǼNjȖOǼȖNjsʰ ¯ȖŘOǼÞŸŘÞض Ř_ ǼsŎƼŸNjĶ _ɴŘŎÞOǣ Ÿ¯ ¯ÞǣÌ ǣǣsŎEͶsǣ ÞŘÌEÞǼÞض OŸǣǼĶ Ķ¶ŸŸŘǣ ʹrĶĶÞŸǼǼ ˓ ^sɟÞĶĶɴˠ˨˨ˤʱəÞʊʊÞŘÞ˓ōʊʊŸĶˡ˟˟ˢʱ®NjŘOŸsǼĶʳˡ˟˟˥ʱōŘʊŸˡ˟ˠ˟ʱ®NjŘʊŸÞʰ ®NjŘOŸ˓ǻŸNjNjÞOsĶĶÞˡ˟ˠ˟ʺŘ__sƼÞOǼs_ǼÌsÞNjĶÞ¯sÌÞǣǼŸNjɴǼNjÞǼǣ ʹ®NjŘOŸsǼĶʳˡ˟ˠˡʱ NɚNjNjŸsǼĶʳˡ˟ˠˣʱDNjÞ¶ŸĶÞŘsǼĶʳˡ˟ˠ˥ʺʳ ǢŸŎsȖǼÌŸNjǣƼNjŸƼŸǣs_¯ȖŘOǼÞŸŘͶȖÞĶ_ƼƼNjŸOÌǣǼŸŸĶǼŸ_sǣONjÞEsǼÌsĶÞ¯s ÌÞǣǼŸNjɴ ǼNjÞǼǣ Ÿ¯ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ Ÿ¯ ǣȖOÌ ǼNjŘǣÞǼÞŸŘĶ sŘɚÞNjŸŘŎsŘǼǣ ʹ®NjŘOŸsǼ Ķʳ ˡ˟˟˧ʺʳÝŘƼNjǼÞOȖĶNjʰʹ®NjŘOŸsǼĶʳˡ˟˟˧ʺƼƼĶÞs_ǼÌs¶ȖÞĶ_ƼƼNjŸOÌǼŸ_sƼÞOǼǼÌs ĶÞŘĨ EsǼɠssŘ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣ Ř_ ǼÌs ȖǼÞĶÞʊǼÞŸŘ Ÿ¯ OŸǣǼĶ ǣɴǣǼsŎǣʰ ɠÌsNjs ǼÌs ¶ȖÞĶ_ Þǣ _s¯ÞŘs_ ǣ ¶NjŸȖƼ Ÿ¯ ǣƼsOÞsǣ ǼÌǼ ǣÌNjs ǼÌsÞNj ŘÞOÌs NjsLJȖÞNjsŎsŘǼǣʰ sɮƼĶŸÞǼÞض ǼÌs ǣŎs Njضs Ÿ¯ sŘɚÞNjŸŘŎsŘǼĶ NjsǣŸȖNjOsǣ ɠÞǼÌ ǣÞŎÞĶNj ÌEÞǼǣ ʹNJŸŸǼ ˠ˨˥˦ʺʳ ŎŸŘ¶ ǼÌs ÞŘǼsNjOǼÞŸŘǣ ǼÌǼ NjsĶǼs ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘ ǼŸ OsNjǼÞŘ sŘɚÞNjŸŘŎsŘǼʰ ǼNjŸƼÌÞONjsĶǼÞŸŘǣÌÞƼǣNjsŸ¯ONjȖOÞĶÞŎƼŸNjǼŘOsŘ_NjsȖǣs¯ȖĶǼŸŸĶǣǼŸǣǣsǣǣǼÌs ¯ȖŘOǼÞŸŘÞضŘ_ǼÌsĶsɚsĶŸ¯OŸŎƼĶsɮÞǼɴŸ¯ǣɴǣǼsŎʳ
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ǢÞŘOsÌEÞǼǼÌsǼsNjŸ¶sŘsÞǼɴÌǣEssŘ_sŎŸŘǣǼNjǼs_ǼŸǣÌƼsǼÌsĶÞ¯sÌÞǣǼŸNjɴǼNjÞǼǣ Ÿ¯ ¯ÞǣÌ ǣƼsOÞsǣ ƼŸƼȖĶǼÞŸŘǣ ʹDNjÞ¶ŸĶÞŘsǼ Ķʳ ˡ˟ˠ˥ʺʰ ɠs ƼNjs_ÞOǼ ǼÌǼ ǼÌÞǣ OŘ ONjsǼs ǣŸŎsɚNjÞǼÞŸŘÞŘǼNjŸƼÌÞOǣǼNjȖOǼȖNjsŸ¯ƼŸƼȖĶǼÞŸŘǣŘ_OŸŘǣsLJȖsŘǼĶɴOŸŎŎȖŘÞǼÞsǣʰ ǼÌǼŎȖǣǼEssɚĶȖEĶsEɴ_sǣONjÞƼǼŸNjǣŸ¯ǼNjŸƼÌÞOŘÞOÌsʳ
ǻÌs ŘĶɴǣÞǣ Ÿ¯ ONjEŸŘ Ř_ ŘÞǼNjŸ¶sŘ ǣǼEĶs ÞǣŸǼŸƼsǣ Ìǣ EssŘ ɠÞ_sĶɴ Ȗǣs_ ÞŘ ǣǼȖ_Þsǣ Ÿ¯ ǼNjŸƼÌÞO sOŸĶŸ¶ɴ ʹĵɴŎŘsǼ Ķʳ ˡ˟ˠˡʺ Ř_ ǼÌs NjsOsŘǼ _sɚsĶŸƼŎsŘǼ Ÿ¯ _sǣONjÞƼǼŸNjǣ ʹƼŸƼȖĶǼÞŸŘˀOŸŎŎȖŘÞǼɴ˚ɠÞ_s ŎsǼNjÞOǣʺ Ř_ ǣǼǼÞǣǼÞOĶ ŎŸ_sĶǣ ʹŎÞɮÞض ŎŸ_sĶʺEǣs_ŸŘǼÌsOŸŘOsƼǼŸ¯ÞǣŸǼŸƼÞOŘÞOÌsÌɚssŎsNj¶s_ǣŎÞŘƼƼNjŸOÌsǣ ǼŸ_sǣONjÞEsLJȖŘǼÞǼǼÞɚsĶɴǼÌsǼNjŸƼÌÞOŘÞOÌsŸ¯ƼŸƼȖĶǼÞŸŘǣŘ_OŸŎŎȖŘÞǼÞsǣŘ_ǼŸ ÞŘ¯sNj ǼÌs ƼǼÌɠɴǣ Ÿ¯ŸNj¶ŘÞO ŎǼǼsNj ÞŘ ɠÞ_s ɚNjÞsǼɴ Ÿ¯ ǣɴǣǼsŎǣ ʹĵɴŎŘsǼĶʳ ˡ˟˟˦ʱğOĨǣŸŘsǼĶʳˡ˟ˠˠʺʳ
NJsǣÞ_sŘǼ¯ÞǣÌƼŸƼȖĶǼÞŸŘǣǼÌǼÞŘÌEÞǼɠÞ_sɚNjÞsǼɴŸ¯sŘɚÞNjŸŘŎsŘǼǣOŘEsȖǣs_ ǣ ǣȖÞǼEĶs ŎŸ_sĶǣ ǼŸ ǣǣsǣǣ ǼÌs ÞŘ¯ĶȖsŘOs Ÿ¯ _Þ¯¯sNjsŘǼ ÌEÞǼǼ ǼɴƼsǣ ŸŘ ǼÌs ɚNjÞEÞĶÞǼɴŸ¯ǼÌsǼNjŸƼÌÞOŘÞOÌsʰǼÞŘǼNjŘ_ÞŘǼsNj˚ͶŸŸŘĶsɚsĶʳ
ÞŎ Ÿ¯ ǼÌÞǣ ɠŸNjĨ Þǣ ǼŸ ǣǣsǣǣ ǼÌs ǼNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ NjsǣÞ_sŘǼ ¯ÞǣÌ ƼŸƼȖĶǼÞŸŘǣONjŸǣǣÌEÞǼǼǣǼÞŘǼsNj˚Ř_ÞŘǼNj˚ͶŸŸŘĶsɚsĶʰɠÞǼÌƼNjǼÞOȖĶNj¯ŸOȖǣ ǼŸǼNj¶sǼǣƼsOÞsǣŸ¯ÌÞ¶ÌsOŸĶŸ¶ÞOĶɚĶȖsʳǻÌsƼȖNjƼŸǣsÞǣǼŸÞŘɚsǣǼÞ¶ǼsÌŸɠǼÌs _Þ¯¯sNjsŘǼ ÌEÞǼǼǣ Ÿ¯ ǼNjŘǣÞǼÞŸŘĶ sOŸǣɴǣǼsŎ ǣÌƼs ǼÌs OŸŎƼŸǣÞǼÞŸŘ Ř_ǼNjŸƼÌÞO ǣǼNjȖOǼȖNjsŸ¯NjsǣÞ_sŘǼ¯ÞǣÌƼŸƼȖĶǼÞŸŘǣʳ
ǢƼsOÞĶǼǼsŘǼÞŸŘɠǣ¶ÞɚsŘǼŸŸŘsŸ¯ǼÌsNjsǣÞ_sŘǼ¯ÞǣÌǣƼsOÞsǣǼÌǼsɮÌÞEÞǼǣɚsNjɴ ÌÞ¶Ì sOŸĶŸ¶ÞOĶ ɚĶȖsʰ ǣ ÞǼ Þǣ EĶs ǼŸ ƼŸƼȖĶǼs ɠÞ_s ɚNjÞsǼɴ Ÿ¯ ÌEÞǼǼǣ ɠÞǼÌÞŘ ŎŘɴ ōs_ÞǼsNjNjŘsŘ OŸǣǼĶ Ķ¶ŸŸŘǣʲ ǼÌs ōs_ÞǼsNjNjŘsŘ ǼŸŸǼÌONjƼ ƼÌŘÞȖǣ ¯ǣOÞǼȖǣʹəĶsŘOÞsŘŘsǣʰˠ˧ˡˠʺʹĵsŸŘNj_Ÿǣ˓ƻsǼNjÞ_Þǣˠ˨˨˥ʱNÞŎŎNjȖǼsǼĶʳˡ˟˟ˢʱ ǻÞ¶ŘŸsǼĶʳˡ˟˟˥ʱNɚNjNjŸsǼĶʳˡ˟ˠˠʺʰĶǣŸOĶĶs_ĨÞĶĶÞ¯Þǣ̯ŸNjÞǼǣǣŎĶĶǣÞʊsʰǣÌŸNjǼ ĶÞ¯sOɴOĶsŘ_ƼsNjǣÞǣǼsŘOɴÞŘǣŎĶĶENjOĨÞǣÌɠǼsNjEŸ_ÞsǣʳÝǼÞǣǼÌsŸŘĶɴǣƼsOÞsǣŸ¯ NɴƼNjÞŘŸ_ŸŘǼÞ_s¯ŎÞĶɴÞŘÌEÞǼÞضÝǼĶÞŘOŸǣǼĶɠǼsNjǣŘ_ǣĶsɚsĶŸ¯ƼNjŸǼsOǼÞŸŘ ÞǼ Þǣ ÞŘOĶȖ_s_ ÞŘ ǼÌs ŘŘsɮ ÝÝ Ÿ¯ ËEÞǼǼǣ ^ÞNjsOǼÞɚs ʹ˨ˡˀˣˢˀrrNʺʰ Njs¶Nj_Þض ǼÌs OŸŘǣsNjɚǼÞŸŘŸ¯ŘǼȖNjĶÌEÞǼǼǣŘ_ɠÞĶ_ ¯ȖŘŘ_¯ĶŸNjÞŘǼÌsrȖNjŸƼsŘȕŘÞŸŘʰ
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Ř_ __ÞǼÞŸŘĶĶɴ ÞǼ Þǣ ĶǣŸ ÞŘOĶȖ_s_ ÞŘ ǼÌs NJs_ ĶÞǣǼ Ÿ¯ ǼÌNjsǼsŘs_ ǣƼsOÞsǣ Ÿ¯ ÝȕNŗ ʹÝŘǼsNjŘǼÞŸŘĶ ȕŘÞŸŘ ¯ŸNj NŸŘǣsNjɚǼÞŸŘ Ÿ¯ ŗǼȖNjsʺʰ ɠÌŸǣs ǼÌNjsǼ Þǣ OĶǣǣÞ¯Þs_ Ÿ¯ ˆĵsǣǼNŸŘOsNjءʳ
ǻɠŸOŸǣǼĶͶŸŸŘǣɠsNjsǼĨsŘÞŘǼŸOOŸȖŘǼʰɠÞǼÌÞŘɠÌÞOÌŘȖŎEsNjŸ¯ÌEÞǼǼǣ ɠsNjs ǣsĶsOǼs_ʰ sÞǼÌsNj ŘǼȖNjĶ ŸNj NjǼÞ¯ÞOÞĶʰ OÌNjOǼsNjÞʊs_ Eɴ _Þ¯¯sNjsŘǼ ĶsɚsĶ Ÿ¯ OŸŘ¯ÞŘsŎsŘǼ Ř_ ɚÞĶEÞĶÞǼɴ Ÿ¯ EǣĶ ǣŸȖNjOsǣʳ ®ȖNjǼÌsNjŎŸNjsʰ ǼÌs OŸŎƼȖǼÞض Ÿ¯ DɴsǣÞŘ ŎÞɮÞض ŎŸ_sĶǣ ĶĶŸɠs_ ǼŸ sǣǼÞŎǼs ǼÌs ɚNjÞEÞĶÞǼɴ Ÿ¯ EǣĶ ǣŸȖNjOsǣ OŸŘǼNjÞEȖǼÞŸŘǼŸǼÌsǼNjŸƼÌÞOƼǼÌɠɴǣǼÌǼǣȖǣǼÞŘǼÌsǼNj¶sǼ¯ÞǣÌONjŸǣǣÌEÞǼǼǣʳ
11 ˠʳˡʳ ōǼsNjÞĶǣŘ_ŎsǼÌŸ_ǣ ǢǼȖ_ɴNjsǣ ǻÌÞǣǣǼȖ_ɴɠǣONjNjÞs_ŸȖǼÞŘǼɠŸĶ¶ŸŸŘsOŸǣɴǣǼsŎǣʲǼÌsǢǼ¶ŘŸŘs_ÞōNjǣĶŘ_ ǼÌsəsŘÞOsĵ¶ŸŸŘÞŘǣŸȖǼÌsNjŘŘ_ŘŸNjǼÌsNjŘÝǼĶɴNjsǣƼsOǼÞɚsĶɴʳ ǻÌs ǢǼ¶ŘŸŘs _Þ ōNjǣĶ ʹˢ˦̨ ˤˡ˂ ŗʰ ˠˡ̨ ˡ˧˂ rʺ Þǣ ǣÌĶĶŸɠ ǣsŎÞ˚sŘOĶŸǣs_ EǣÞŘ ɠÞǼÌǣȖNj¯OsNjsŸ¯ƼƼNjŸɮÞŎǼsĶɴˡˠĨŎˡŘ_ŘɚsNj¶s_sƼǼÌŸ¯ˠŎʹ®Þ¶ʳˠʺʳ ŗŸ ǣȖEǣǼŘǼÞĶ ÞŘƼȖǼǣ Ÿ¯ ¯NjsǣÌɠǼsNj Njs ƼNjsǣsŘǼʰ ǼÌsNjs¯ŸNjs ɚĶȖsǣ Ÿ¯ ǼsŎƼsNjǼȖNjs Ř_ǣĶÞŘÞǼɴNjs¶sŘsNjĶĶɴÌÞ¶ÌsNjÞŘOŸŎƼNjÞǣŸŘǼŸǼÌs_ĠOsŘǼǣsʹōʊʊŸĶsǼĶʳ ˡ˟ˠ˟ʺʰ ǼŸ ɠÌÞOÌ ǼÌs ǼɠŸ ŎÞŘ ǣȖE˚EǣÞŘǣ Ìɚs _Þ¯¯sNjsŘǼ _s¶Njss Ÿ¯ OŸŘŘsOǼÞŸŘʳ ǻÌsŘŸNjǼÌsNjŘǣȖE˚EǣÞŘÌǣŘNjNjŸɠŘ_ǣÌĶĶŸɠOÌŘŘsĶʹˣˤ˟ŎɠÞ_ǼÌŘ_ˠ˟˚ˢ˟ OŎ _sƼǼÌʺ ǼÌǼ ĶĶŸɠǣ ŸŘĶɴ ǣONjOs ɠǼsNj sɮOÌضsʰ ǼÌȖǣ NjsƼNjsǣsŘǼǣ ǼÌs OǼȖĶ Ķ¶ŸŸŘ EǣÞŘʰ ɠÞǼÌ ÌÞ¶ÌsNj ŘŘȖĶ ɚNjÞEÞĶÞǼɴ Ÿ¯ ǼÌs ŎÞŘ ƼÌɴǣÞOŸ˚OÌsŎÞOĶ ƼNjŎsǼsNjǣ Ř_ ǼÌs EŸǼǼŸŎ Þǣ Ÿ¯ǼsŘ _ŸŎÞŘǼs_ Eɴ sɚsŘǼǣ Ÿ¯ ǣs_ÞŎsŘǼ NjsǣȖǣƼsŘǣÞŸŘʳDɴOŸŘǼNjǣǼʰǼÌsǣŸȖǼÌsNjŘǣȖE˚EǣÞŘÞǣOŸŘŘsOǼs_ǼŸǼÌsǣsǼÌNjŸȖ¶Ì ɠÞ_sNjŎŸȖǼÌʹˠˢˤ˟ŎɠÞ_ǼÌŘ_EŸȖǼˡŎ_sƼǼÌʺǼÌǼÞŘ_ȖOsǣÌÞ¶ÌsNjNjǼsŸ¯ ɠǼsNjǼȖNjŘŸɚsNjʹĵĵŸ¶¶ÞsǼĶʳˡ˟˟ˣʺŘ_ǣǼNjŸŘ¶sNjŎNjÞŘsÞŘ¯ĶȖsŘOsʳǻÌsǢǼ¶ŘŸŘs Þǣ ĶǣŸ OĶǣǣÞ¯Þs_ ǣ ŸĶÞ¶ŸǼNjŸƼÌÞOʰ ǣ ǼÌs ŎsŘ OÌĶŸNjŸƼÌɴĶĶ˚ OŸŘOsŘǼNjǼÞŸŘ Þǣ NjŸȖŘ_ˠʳ˟ͼ¶Ķ˚ˠÞŘǼÌsɠǼsNjOŸĶȖŎŘʹǢNjʰĵsŸŘNj_Þ˓ōʊʊŸĶˠ˨˨˨ʺŘ_ˢʳ˟ͼ¶
Ķ˚ˠ ÞŘ ǼÌs ǣs_ÞŎsŘǼ ʹƻȖǣOs__ȖsǼ Ķʳ ˠ˨˨˨ʺʳ OOŸNj_Þض ǼŸ ǼÌs rȖNjŸƼsŘ ^ÞNjsOǼÞɚs
˨ˡˀˣˢˀNrrʰ ǼÌs Njs Þǣ ÞŘOĶȖ_s_ ÞŘ ǼÌs ĶÞǣǼǣ Ÿ¯ ǼÌs ǢNÝ ʹÝǻ˟ˠ˟˟ˡ˥ʺ Ř_ Ǣƻ ʹÝǻ˟ˠ˟˟ˡ˧ʺʳ ǻÌsͶŸŸŘŸ¯əsŘÞOsʹˣˤ̨ˡ˥˅ŗʰˠˡ̨ˡ˟˅rʺÞǣǼÌsĶNj¶sǣǼͶŸŸŘĶǣɴǣǼsŎÞŘrȖNjŸƼs ʹ®Þ¶ʳˠʺʰÌǣǣȖNj¯OsNjsŸ¯EŸȖǼˤˤ˟ĨŎˡŘ_OOŸNj_ÞضǼŸÞǼǣÌɴ_NjŸĶŸ¶ɴOŘEs _ÞɚÞ_s_ ÞŘ ǼÌNjss ŎÞŘ ǣȖE˚EǣÞŘǣʲ ŗŸNjǼÌsNjŘʰ NsŘǼNjĶ Ř_ ǢŸȖǼÌsNjŘ ʹɚŘʊÞsǼ Ķʳ ˠ˨˦˨ʺʳÝǼÌǣŎÞŘĶɴǣÌĶĶŸɠɠǼsNjǣʹŎsŘ_sƼǼÌEŸȖǼˠ̝˟ʳˢŎʰDŸŘ¯sǼĶʳˡ˟˟ˣʺ ONjŸǣǣs_Eɴ_ssƼsNjǣÌÞƼƼÞضOÌŘŘsĶǣsɮǼsŘ_ÞضǼŸɠNj_ǣǼÌsǼÌNjssŎŸȖǼÌǣʹĵÞ_Ÿʰ ōĶŎŸOOŸ Ř_ NÌÞŸ¶¶Þʺ ǼÌǼ ĶĶŸɠ OŸŘŘsOǼÞŸŘ Ř_ ɠǼsNj sɮOÌضs ɠÞǼÌ ǼÌs
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_NjÞǼÞO Ǣsʳ ǻÌs ŎĠŸNj ¯NjsǣÌɠǼsNj ǼNjÞEȖǼNjÞsǣ Njs ĶŸOǼs_ ÞŘ ǼÌs ŘŸNjǼÌsNjŘ ǣȖE˚ EǣÞŘʰÌsŘOsÌǣĶŸɠsNjǣĶÞŘÞǼɴǼÌŘǼÌsŸǼÌsNjǣȖE˚EǣÞŘǣʳǻÌsNjsÞǣǣȖEĠsOǼs_ǼŸ ǼÌsɠÞ_sǣǼǼÞ_ĶNjضsŸ¯ǼÌsōs_ÞǼsNjNjŘsŘǢsʰɠÌÞOÌÞǣȖƼǼŸƼƼNjŸɮÞŎǼsĶɴˠŎʳ ǻÌs ÌÞ¶Ì ɚNjÞEÞĶÞǼɴ Ÿ¯ ŎŸNjƼÌŸĶŸ¶ÞOĶ Ř_ ƼÌɴǣÞOŸ˚OÌsŎÞOĶ ƼNjŎsǼsNjǣ ÞŘ_ȖOsǣ ÌÞ¶Ì ĶsɚsĶǣ Ÿ¯ ǣƼǼÞĶ Ř_ ÌEÞǼǼ ÌsǼsNjŸ¶sŘsÞǼɴ ǼÌǼ OÌNjOǼsNjÞʊsǣ ǼÌs ɠÌŸĶs ͶŸŸŘʳŎŸŘ¶ǼÌs_Þ¯¯sNjsŘǼĶsɚsĶǣŸ¯ƼNjŸǼsOǼÞŸŘŸ¯ǼÌsNjsʰǼÌsËEÞǼǼǣ^ÞNjsOǼÞɚs ʹ˨ˡˀˣˢˀNrrʺÞŘOĶȖ_sǣǼÌsŗŸNjǼÌsNjŘĵ¶ŸŸŘŸ¯əsŘÞOsɠÞǼÌÞŘǼÌsǢÞǼsǣŸ¯NŸŎŎȖŘÞǼɴ ÝŎƼŸNjǼŘOsʹǢNÝʺǣɠsĶĶǣǼÌsNsŘǼNjĶ˚ǢŸȖǼÌsNjŘNjsǣʹNjsǣƼsOǼÞɚsĶɴÝǻˢˡˤ˟˟ˢ˟Ř_ Ýǻˢˡˤ˟˟ˢˠʺʰŘ_ǼÌsͶŸŸŘǣɠÌŸĶsÞǣ_sǣÞ¶ŘǼs_ǣǢƼsOÞĶƻNjŸǼsOǼÞŸŘNjs ʹǢƻʰÝǻˢˡˤ˟˟ˣ˥ʺʳ ǢŎƼĶÞضǣÞǼsǣ ®ÞɚsǣÞǼsǣOŸNjNjsǣƼŸŘ_ÞضǼŸ_Þ¯¯sNjsŘǼŘǼȖNjĶŘ_NjǼÞ¯ÞOÞĶÌEÞǼǼǣʰɠÌsNjsǼÌsǼNj¶sǼ ǣƼsOÞsǣÞǣɠÞ_sĶɴ_ÞǣǼNjÞEȖǼs_ʰɠsNjsOÌŸǣsŘONjŸǣǣǼÌsǼɠŸǣǼȖ_ɴNjsǣʹ®Þ¶ʳˠʺʳ ®Þ¶ȖNjsˠʳĵŸOǼÞŸŘŸ¯ǼÌsǣǼȖ_ɴNjsǣŘ_NjsĶǼÞɚsƼŸǣÞǼÞŸŘŸ¯ǼÌsÌEÞǼǼǣsɮŎÞŘs_ʳ
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ÝŘǼÌsǢǼ¶ŘŸŘs_ÞōNjǣĶǼÌs¯ŸĶĶŸɠÞضǼɠŸǣÞǼsǣɠsNjsÞŘɚsǣǼÞ¶Ǽs_ʲ
Ǣµʲ Ǣs¶Njǣǣ Ŏs_Ÿɠ ˛ Þǣ _sŘǣsNɴŎŸ_ŸOs ŘŸ_Ÿǣ ʹȕONjÞ ǣOÌsNjǣŸŘʰ ˠ˧˦˟ʺ Ŏs_Ÿɠ ĶŸOǼs_ ŘsNj ƻȖŘǼ _˅Ͷʰ ÞŘ ǼÌs ǣŸȖǼÌsNjŘ ǣȖE˚EǣÞŘ Ÿ¯ ǼÌs ǢǼ¶ŘŸŘsʳǢÞǼs_sƼǼÌÞǣEŸȖǼˣ˟OŎŘ_ǼÌsǣs_ÞŎsŘǼÞǣǣŘ_ɴ˚ƼsĶÞǼÞOʰ¯ȖĶĶɴ OŸɚsNjs_ÞŘNʳŘŸ_ŸǣŘ_ŎÞŘŸNjŎONjŸĶ¶sʳ rƻʲrɚƼŸNjǼÞŸŘƼŸŘ_˛ĶǣŸOĶĶs_ˆOŸŸĶÞضɚǼˇʰÞǣŸŘsŸ¯ǼÌsǼŘĨǣŸ¯ǼÌsǣĶǼɠŸNjĨ ǣɴǣǼsŎĶŸOǼs_ÞŘǼÌsOsŘǼNjĶNjsŸ¯ǼÌsǢǼ¶ŘŸŘsʳǻÌsǼŘĨÞǣƼƼNjŸɮÞŎǼsĶɴ ˥˟ʳ˟˟˟ŎˡÞŘǣȖNj¯OsNjsŘ_ˤ˟OŎ_ssƼʰÞǼÞǣsŘOĶŸǣs_¯NjŸŎǼÌs_ĠOsŘǼ ǣŸȖŘ_Ř_ʰŸŘĶɴŸOOǣÞŸŘĶĶɴʰǣŎĶĶOÌŘŘsĶÞǣŸƼsŘs_ǼŸĶĶŸɠǼÌsÞŘOŸŎs Ÿ¯ɠǼsNj¯ŸNjŘsɠǣĶǼƼNjŸ_ȖOǼÞŸŘʳǢŘ_ɴ˚ƼsĶÞǼÞOEŸǼǼŸŎÞŘOŸɚsNjs_EɴŎÞɮs_ ƼǼOÌsǣŸ¯NʳŘŸ_ŸǣŘ_ŎONjŸĶ¶sʳ ÝŘəsŘÞOsͶŸŸŘǼÌs¯ŸĶĶŸɠÞضǼÌNjssǣÞǼsǣɠsNjsÞŘɚsǣǼÞ¶Ǽs_ʲ Ǣōˠʲ ǢĶǼ ŎNjǣÌ ˠ ˛ Þǣ ǣŎĶĶ ǣĶǼ ŎNjǣÌ ǣÞǼȖǼs_ ÞŘ DOOŘ ʹˣˤʳˣˣ˦ˣˠ̨ŗʰ ˠˡʳˣˠˤ˧ˢ̨rʺʰǣÞǼsOĶŸǣsǼŸǼÌsŘŸNjǼÌsNjŘŎŸȖǼÌŸ¯ǼÌsͶŸŸŘʳ^sƼǼÌÞǣEŸȖǼ ˢ˟OŎʰǼÌsǣs_ÞŎsŘǼÞǣŎȖ__ɴŘ_ÌǣĶŸɠOŸɚsNj¶sʹˤ˚ˡˤ̇ʺŸ¯ǣs¶Njǣǣsǣ ɠÞǼÌǣƼNjǣs_NjÞ¯ǼŎONjŸĶ¶sʳ ǢōˡʲǢĶǼŎNjǣÌˡ˛ÞǣǣĶǼŎNjǣÌƼĶOs_ÞŘŘÞŘŘsNjNjsŸ¯ǼÌsͶŸŸŘʰÞŘǼÌs ƼNjŸɮÞŎÞǼɴŸ¯NNjsɚŘÞǣĶŘ_ʹˣˤʳˣ˦ˠ˨˟̨ŗʰˠˡʳˣˢ˟˧˟̨rʺʳǻÌsǣÞǼsÞǣˣ˟OŎ_ssƼ Ř_ ǼÌs ǣȖEǣǼNjǼs Þǣ ŎÞŘĶɴ ŎȖ__ɴ Ř_ ENjsʰ ɠÞǼÌ ɚsNjɴ ĶŸɠ ʹˤ˚ˠ˟̇ʺ ƼNjsǣsŘOs Ÿ¯ ǣs¶Njǣǣsǣ Ř_ ŎONjŸĶ¶sʳ ÝǼ Þǣ ĶǣŸ OÌNjOǼsNjÞʊs_ Eɴ ǼÌs ƼNjsǣsŘOsŸ¯ǣŎĶĶÞŘǼsNjǼÞ_ĶONjssĨǣʰEŸȖǼˠ˟OŎ_ssƼʳ ^ʲNjǼÞ¯ÞOÞĶ_ÞǼOÌ˛ÞǣŘNjǼÞ¯ÞOÞĶǣɴǣǼsŎŸ¯ǣŎĶĶ_ÞǼOÌsǣǣÞǼȖǼs_ÞŘəÞ¶ŘŸĶsÞǣĶŘ_ ʹˣˤʳˣˣˡˤ˨̨ŗ ˠˡʳˢ˦ˤˢ˧̨rʺʰȖǣs_ÞŘǼÌsƼǣǼǣÞŘĶŘ_¯ÞǣÌƼŸŘ_ʳŗŸɠ_ÞǼOÌsǣ NjsEŸȖǼ˥˟OŎ_ssƼŘ_NjsÞǣŸĶǼs_¯NjŸŎǼÌsNjsǣǼŸ¯ǼÌsͶŸŸŘʳ
14 ǢŎƼĶsOŸĶĶsOǼÞŸŘ ®ÞǣÌOŸŎŎȖŘÞǼÞsǣŘ_ĶĶǼÌsƼŸǣǣÞEĶsǣŸȖNjOsǣŸ¯ŸNj¶ŘÞOŎǼǼsNjNjsƼNjsǣsŘǼǼÞɚsŸ¯ ǼÌs ÌEÞǼǼǣ sɮŎÞŘs_ ɠsNjs OŸĶĶsOǼs_ ÞŘ ǣƼNjÞض Ř_ ȖǼȖŎŘ ˡ˟ˠˣʰ ¯ŸĶĶŸɠÞض ǣǼŘ_Nj_Þʊs_ƼNjŸǼŸOŸĶȖǣs_ÞŘĶĶǼÌsǣǼȖ_ɴǣÞǼsǣʳ ®ÞǣÌɠsNjsOŸĶĶsOǼs_EɴǣsŎÞ˚LJȖŘǼÞǼǼÞɚsOǼOÌʰȖǣÞضǣŎĶĶʹˣ˚˧˚ˠ˟ŎʺEsOÌǣsÞŘs ŘsǼǣɠÞǼÌŎsǣÌǣÞʊsŸ¯ˡŎŎʰOŸɚsNjÞضǼɠŸǼÞŎsǣĨŘŸɠŘǣȖNj¯OsNjsʳǢÞʊsŸ¯ ǼÌs ǣsÞŘs ŘsǼ Ȗǣs_ Ř_ ǣȖNj¯Os Njs ǣŎƼĶs_ ɚNjÞs_ Eɴ ǣǼȖ_ɴ Njs ŸNj ǣÞǼsʰ _sƼsŘ_Þض ŸŘ ǼÌs ǣƼsOÞ¯ÞO ÌEÞǼǼ ÞŘɚŸĶɚs_ ʹsʳ¶ʳ ŘNjNjŸɠsNj ŘsǼ OŸɚsNjÞض ĶŸŘ¶sNj _ÞǣǼŘOs ɠÌsŘ ǣŎƼĶÞض ÞŘ OŘĶǣ ŸNj ONjssĨǣʰ ɠÞ_sNj ŘsǼ ŸɚsNj ɠÞ_sNj ǣȖNj¯Os Njs ɠÌsŘǣŎƼĶÞضÞŘŸƼsŘǣĶǼŎNjǣÌsǣʺʳ ƻNjÞŎNjɴƼNjŸ_ȖOsNjǣǣȖOÌǣǣs¶NjǣǣsǣʰŎONjŸĶ¶sŘ_ǼsNjNjsǣǼNjÞĶÌĶŸƼÌɴǼsǣɠsNjs OŸĶĶsOǼs_ŎŘȖĶĶɴ¯NjŸŎǼÌNjssNjŘ_ŸŎƼŸÞŘǼǣOÌŸǣsŘɠÞǼÌÞŘǼÌsǣŎƼĶs_Njsǣʳ ǢsɠǼsNjǣŎƼĶsǣʹˠ˟ĶŘss_s_ÞŘǼÌsŸĶÞ¶ŸǼNjŸƼÌÞOǢǼ¶ŘŸŘs_ÞōNjǣĶŘ_ˡĶÞŘ əsŘÞOsĵ¶ŸŸŘʺŘ_ǣȖƼsNj¯ÞOÞĶǣs_ÞŎsŘǼOŸNjsǣʹˢOŎƩʺɠsNjsOŸĶĶsOǼs_ÞŘǼNjÞƼĶÞOǼs ǼŸŸEǼÞŘǼÌsǣŸȖNjOsǣNjsĶǼs_NjsǣƼsOǼÞɚsĶɴǼŸǼÌsǣȖǣƼsŘ_s_ƼNjǼÞOȖĶǼsʹƻŷōʺŘ_ ǣs_ÞŎsŘǼNjɴ ŸNj¶ŘÞO ŎǼǼsNj ʹǢŷōʺʳ ÝŘ __ÞǼÞŸŘʰ ŸŘĶɴ ÞŘ əsŘÞOs ĵ¶ŸŸŘ ǣÞǼsǣʰ ʊŸŸƼĶŘĨǼŸŘɠǣǣŎƼĶs_EɴÌŸNjÞʊŸŘǼĶÌȖĶɠÞǼÌǣŎĶĶƼĶŘĨǼŸŘŘsǼɠÞǼÌˠ˥˟ ͼŎ ŎsǣÌ ǣÞʊsʳ ĶĶ ǼÌs ǣŎƼĶsǣ OŸĶĶsOǼs_ ɠsNjs ĨsƼǼ OŸŸĶ ȖŘǼÞĶ NjNjÞɚÞض Ǽ ǼÌs ĶEŸNjǼŸNjɴŘ_ǣǼŸNjs_Ǽ˚ˡ˟̨NƼNjÞŸNjǼŸǼÌsƼNjŸOsǣǣÞضʳ ǢŎƼĶsƼNjŸOsǣǣÞضŘ_ĶEŸNjǼŸNjɴŘĶɴǣÞǣ ®ÞǣÌǣŎƼĶs_ɠsNjsÞ_sŘǼÞ¯Þs_ǼǣƼsOÞsǣĶsɚsĶŘ_ǣsɮʹɠÌsŘƼŸǣǣÞEĶsʺŘ_ǼÌsŎÞŘ EÞŸŎsǼNjÞOǣǣȖOÌǣǣǼŘ_Nj_ĶsضǼÌʹǢĵʺʰǼŸǼĶɠsÞ¶ÌǼʹǻɟʺʰɠsNjsŎsǣȖNjs_ÞŘŸNj_sNj ǼŸ_sǼsNjŎÞŘsǼÌsŎÞŘ_sŎŸ¶NjƼÌÞOǼNjÞǼǣŸ¯ǼÌsƼŸƼȖĶǼÞŸŘǣsɮŎÞŘs_ʳ
¯ǼsNjɠNj_ǣʰ Ǽ ĶsǣǼ ˠˤ ÞŘ_ÞɚÞ_ȖĶǣ ¯ŸNj sOÌ ǣƼsOÞsǣ Ř_ ǣsɮʰ ɠsNjs _ÞǣǣsOǼs_ Eɴ ŎsŘǣ Ÿ¯ ǣOĶƼsĶ Ř_ ǼɠssʊsNjǣ ǼŸ sɮǼNjOǼ ǼÌs _ŸNjǣĶ ŎȖǣOĶsǣʳ Ǣs¶Njǣǣsǣʰ ŎONjŸĶ¶s Ř_ ǼsNjNjsǣǼNjÞĶ ÌĶŸƼÌɴǼsǣ ɠsNjs Þ_sŘǼÞ¯Þs_ Ǽ ǣƼsOÞsǣ ĶsɚsĶʱ sɚsŘǼȖĶ sƼÞƼÌɴǼsǣɠsNjsǣONjƼs_ŎŘȖĶĶɴʳ
15
ɟǼsNj ǣŎƼĶsǣ ɠsNjs ƼNjs˚¯ÞĶǼsNjs_ ʹˡ˟˟ ͼŎʺ Ř_ ǼÌsŘ ¯ÞĶǼsNjs_ ŸŘ ƼNjsOŸŎEȖǣǼs_ ʹˣˤ˟̨NʰˣÌʺ¯ÞĶǼsNjǣʹµ®ˀ®ɟÌǼŎŘʰƼŸNjsǣÞʊs˟ʳ˦ͼŎʺǼŸOŸŘOsŘǼNjǼsǼÌsƻŷōʳǻÌs ǼŸƼ ˠʳˤ OŎ Ÿ¯ ǣs_ÞŎsŘǼ OŸNjsǣ ɠsNjs ǣĶÞOs_ Ř_ ÌŸŎŸ¶sŘÞʊs_ ǣ ƼNjŸɮɴ ¯ŸNj Ǣŷōʳ ʉŸŸƼĶŘĨǼŸŘ ǣŎƼĶsǣ ɠsNjs ONjs¯ȖĶĶɴ OĶsŘs_ ȖŘ_sNj EÞŘŸOȖĶNj ŎÞONjŸǣOŸƼs ǼŸ NjsŎŸɚsŘɴ_sǼNjÞǼĶŎǼsNjÞĶŘ_OŸŘOsŘǼNjǼs_ÞŘsƼƼsŘ_ŸNj¯ǼȖEsǣʳ
ŷŘOs ƼNjŸOsǣǣs_ʰ ĶĶ ǣŎƼĶsǣ ÞŘǼsŘ_s_ ¯ŸNj ǼÌs ŘĶɴǣÞǣ Ÿ¯ ǣǼEĶs ÞǣŸǼŸƼsǣ ɠsNjs ŸɚsŘ˚_NjÞs_Ǽ˥˟̨NǼŸOŸŘǣǼŘǼɠsÞ¶ÌǼ¯ŸNjEŸȖǼˣ˧ÌŘ_ǼÌsŘ ¶NjŸȖŘ_ǼŸ¯ÞŘs ƼŸɠ_sNjȖǣÞضŎÞONjŸŎÞĶĶŸNjŎŸNjǼNjŘ_ƼsǣǼĶsʳƻNjǼŸ¯ǼÌsǣŎƼĶsǣOŸŘǼÞŘÞض ŘɴONjEŸŘǼsOŸŎƼŸŘsŘǼǼÌǼŎÞ¶ÌǼÞŘǼsNj¯sNjsɠÞǼÌǼÌs_sǼsNjŎÞŘǼÞŸŘŸ¯ONjEŸŘ ÞǣŸǼŸƼÞO ǣÞ¶ŘǼȖNjs ɠǣ OÞ_Þ¯Þs_ ʹËNĶ ˠōʺʳ ĵǣǼĶɴʰ Ř ĶÞLJȖŸǼ Ÿ¯ sOÌ ǣŎƼĶs ɠǣ ƼOĨs_ÞŘǼÞŘOƼǣȖĶsǣŘ_ŘĶɴǣs_¯ŸNjδˠˢNŘ_δˠˤŗȖǣÞضŘÞǣŸǼŸƼsNjǼÞŸŎǣǣ
ǣƼsOǼNjŸŎsǼsNj ʹǻÌsNjŎŸ˚rĶsOǼNjŸŘ ^sĶǼ ƻĶȖǣ ɭƻʺ OŸȖƼĶs_ ǼŸ Ř sĶsŎsŘǼĶ ŘĶɴǣsNj ʹǻÌsNjŎŸ˚rĶsOǼNjŸŘ®ĶǣÌrˠˠˠˡʺʳ NNjEŸŘ Ř_ ŘÞǼNjŸ¶sŘ ǣǼEĶs ÞǣŸǼŸƼs NjǼÞŸǣ ɠsNjs sɮƼNjsǣǣs_ ÞŘ δ ȖŘÞǼ ŘŸǼǼÞŸŘʰ ǣ ƼNjǼǣ ƼsNj ŎÞĶ _sɚÞǼÞŸŘ ¯NjŸŎ ǼÌs ÞŘǼsNjŘǼÞŸŘĶ ǣǼŘ_Nj_ǣ ʹNjsǣƼsOǼÞɚsĶɴ ƻss ^ss DsĶsŎŘÞǼs Ř_ ǼŎŸǣƼÌsNjÞO ŗˡ ¯ŸNj ONjEŸŘ Ř_ ŘÞǼNjŸ¶sŘʺ Ř_ _sǼsNjŎÞŘs_ ǣ ¯ŸĶĶŸɠǣʲ δɭ̠ʻʹNJǣŎƼĶsˀNJǣǼŘ_Nj_ʺ̜ˠʼ̞ˠ˟ˢʰ ɠÌsNjs ɭ ÞǣˠˢN ŸNjˠˤŗ Ř_ NJ Þǣ ǼÌs NjsĶǼÞɚs ˠˢNˀˠˡNŸNjˠˤŗˀˠˣŗNjǼÞŸʳ ^ǼŘĶɴǣÞǣ ^sŎŸ¶NjƼÌÞOǼNjÞǼǣ˚ƻNjsĶÞŎÞŘNjɴŘĶɴǣÞǣŸ¯ǼÌs¯Þǣ̯ȖŘ_ǼĶĶŸɠs_ǼŸǣǣsǣǣ ǼÌsOŸŘǼsɮǼŸ¯ǼÌsOŸŎŎȖŘÞǼɴɠÌsNjsǼÌsǼNj¶sǼǣƼsOÞsǣɠǣƼĶOs_ʳ
ǻÌs ŘȖŎEsNj Ÿ¯ ÞŘ_ÞɚÞ_ȖĶǣ Ÿ¯ sOÌ ǣƼsOÞsǣ OŸĶĶsOǼs_ ɠǣ NjsĶǼs_ ǼŸ ǼÌs ǣƼsOÞ¯ÞO ǣŎƼĶÞضNjsŘ_ǼÌsŘǣǼŘ_Nj_Þʊs_ǼŸˠ˟˟ŎˡʰÞŘŸNj_sNjǼŸOŸŎƼNjsǼÌs_sŘǣÞǼɴ
ŎŸŘ¶ǣÞǼsǣǣŘ̨ÞŘ_ÞɚÞ_ȖĶǣˠ˟˟˚ˡŘ_NjsĶǼÞɚsƼsNjOsŘǼ¶sEȖŘ_ŘOsʳÝŘ__ÞǼÞŸŘʰ
sOÌ ǣƼsOÞsǣ ɠǣ ǣǣÞ¶Řs_ ǼŸ Ř sOŸĶŸ¶ÞOĶ Ř_ ¯ss_Þض ¶NjŸȖƼʰ ǣ _s¯ÞŘs_ ÞŘ ōĶɚǣÞsǼ Ķʳ ʹˡ˟˟ˣʺ Ř_ ®NjŘOŸsǼ Ķʳ ʹˡ˟˟˥ʺʰ ǼŸ ŸȖǼĶÞŘs ǼÌs ŎÞŘ sOŸĶŸ¶ÞOĶ ¯sǼȖNjsǣŸ¯ǼÌsƼŸƼȖĶǼÞŸŘǣ¯ŸȖŘ_ɠÞǼÌÞŘǼÌs¯ÞǣÌǣǣsŎEͶsǣʳ
16
ÝŘ ŸNj_sNj ǼŸ ¯ŸOȖǣ ŸŘ ǼÌs ƼŸƼȖĶǼÞŸŘ ĶsɚsĶ Ÿ¯ ǼÌs ǼNj¶sǼ ǣƼsOÞsǣʳ ¯ǣOÞǼȖǣʰ ǼÌs ŎÞŘ ǼNjÞǼǣ Ÿ¯ ǼÌs _sŎŸ¶NjƼÌÞO ǣǼNjȖOǼȖNjs ɠsNjs _sǣONjÞEs_ Eɴ ĶŸŸĨÞض Ǽ ǼÌs ¯NjsLJȖsŘOɴ _ÞǣǼNjÞEȖǼÞŸŘǣ Ÿ¯ ǣǼŘ_Nj_ ĶsضǼÌ ʹǢĵʺ Ř_ ǼÌs ǣsɮ NjǼÞŸ ¯sŎĶsʲŎĶs ʹ®ʲōʺʳÝŘ __ÞǼÞŸŘʰ ǼNjŸƼÌÞO ƼŸǣÞǼÞŸŘǣ ʹǻƻʺ ɠsNjs sǣǼÞŎǼs_ ¯ŸNjʳ ¯ǣOÞǼȖǣ ȖǣÞض ǼÌs sLJȖǼÞŸŘNjsƼŸNjǼs_ÞŘʹƻŸǣǼˡ˟˟ˡʺʲǻƻ̠ʻʹδˠˤŗ ¯˚δˠˤŗEʺˀͶŘʼ̛ʭʰɠÌsNjsδˠˤŗ¯Ř_ δˠˤŗENjsNjsǣƼsOǼÞɚsĶɴǼÌsŘÞǼNjŸ¶sŘÞǣŸǼŸƼÞOǣÞ¶ŘǼȖNjsŸ¯ǼÌs¯ÞǣÌǣƼsOÞsǣŘ_ǼÌǼŸ¯ ǼÌsǣŸȖNjOsEǣsĶÞŘsʳǣǣŸȖNjOsEǣsĶÞŘsɠsȖǣs_ǼÌsŎsŘɚĶȖsŸ¯ĶĶǼÌsǣŎƼĶsǣ Ÿ¯ ʊŸŸƼĶŘĨǼŸŘ OŸĶĶsOǼs_ ɠÞǼÌÞŘ ǼÌs əsŘÞOs ͶŸŸŘ ʹ˦ʳˢ˦̈ʺ Ř_ ǼÌs ǢǼ¶ŘŸŘs _Þ ōNjǣĶʹ˥ʳˤ̈ʺʳͶŘ̠ˢʳˣ̈ÞǣǼÌsǼNjŸƼÌÞOsŘNjÞOÌŎsŘǼsɮƼsOǼs_¯ŸNjsOÌǼNjŸƼÌÞOĶsɚsĶ Ř_ʭÞǣǼÌsǼNjŸƼÌÞOƼŸǣÞǼÞŸŘŸ¯ǼÌsǣƼsOÞsǣȖǣs_ǣEǣsĶÞŘsʰǼÌǼÞǣˡʳ ǻNjŸƼÌÞOǣǼNjȖOǼȖNjs˚^Þ¯¯sNjsŘOsǣ¯ŸNjÞǣŸǼŸƼsǣEsǼɠssŘͶŸŸŘǣŘ_ŎŸŘ¶ÌEÞǼǼǣ ɠsNjsOŸŘ_ȖOǼs_ǣsƼNjǼsĶɴ¯ŸNjδˠˢNŘ_δˠˤŗʰNjsǣƼsOǼÞɚsĶɴǼÌNjŸȖ¶ÌǼ˚ǼsǣǼŸNjŘŸŘ˚ ƼNjŎsǼNjÞOħNjȖǣĨĶ˚ɟĶĶÞǣ¯ǼsNjǼsǣǼÞض¯ŸNjŘŸNjŎĶ_ÞǣǼNjÞEȖǼÞŸŘŸ¯ǼÌs_Ǽʳ
ÝŘ ŸNj_sNj ǼŸ ǣǣsǣǣ ǼÌs ÞǣŸǼŸƼÞO ŘÞOÌs ɠÞ_ǼÌ Ř_ NjsĶǼÞɚs ŎsǼNjÞOǣʰ ǣǼŘ_Nj_ sĶĶÞƼǣs NjsǣʹǢrOʰOŸNjNjsOǼs_¯ŸNjǣŎĶĶǣŎƼĶsǣÞʊsʺɠsNjssǣǼÞŎǼs_EɴDɴsǣÞŘǣǼǼÞǣǼÞOǣ ȖǣÞضǼÌsNJƼOĨ¶sǢÝDrNJɚˡʳ˟ʳˡʹǢǼEĶsÝǣŸǼŸƼsDɴsǣÞŘrĶĶÞƼǣsǣÞŘNJʺʹğOĨǣŸŘsǼ Ķʳˡ˟ˠˠʺŘ_OŸŎŎȖŘÞǼɴ˚ɠÞ_sŎsǼNjÞOǣɠsNjsOĶOȖĶǼs_ȖǣÞضǼÌsNJƼOĨ¶sǢÝNJ ˣʳˡʳˡʹƻNjŘsĶĶsǼĶʳˡ˟ˠ˟ʺʳ ōsǼNjÞOǣ ɠsNjs sǣǼÞŎǼs_ ÞŘ_sƼsŘ_sŘǼĶɴʰ Ǽ ¯ÞNjǣǼʰ ¯ŸNj ĶĶ Ÿ¯ ǼÌs ǣƼsOÞsǣ ƼŸƼȖĶǼÞŸŘǣ ¯ŸȖŘ_ÞŘĶĶŸ¯ǼÌsÌEÞǼǼǣŘ_¯ŸNjEŸǼÌǣsǣŸŘǣʰÞŘŸNj_sNjǼŸÌɚsŘŸɚsNjĶĶɚÞsɠ Ÿ¯ǼÌs¯ÞǣÌOŸŎŎȖŘÞǼɴŘ_ǼÌsɚNjÞEÞĶÞǼɴÞŘǼÞŎsʳǢsOŸŘ_ĶɴʰǼŸ¯ŸOȖǣŸŘǼÌsǼNj¶sǼ ǣƼsOÞsǣ Ř_ ǣǣsǣǣ ÞǼǣ ǼNjŸƼÌÞO ŘÞOÌs ɚNjÞEÞĶÞǼɴ ONjŸǣǣ ÌEÞǼǼǣʰ ǼÌs ǣŎs ŎsǼNjÞOǣ ɠsNjssǣǼÞŎǼs_ŸŘĶɴ¯ŸNjʳ¯ǣOÞǼȖǣƼŸƼȖĶǼÞŸŘǣʰƼŸŸĶÞضǣsǣŸŘǣǼŸ¶sǼÌsNjʳ
ŎŸŘ¶ǼÌsOŸŎŎȖŘÞǼɴ˚ɠÞ_sŎsǼNjÞOʰÞŘƼNjǼÞOȖĶNjǼÌs¯ŸĶĶŸɠÞضǣɠsNjsOĶOȖĶǼs_¯ŸNj ǼÌsƼȖNjƼŸǣsʲ
i) δˠˤŗ NJضs ʹŗNJʺʰ _Þ¯¯sNjsŘOs EsǼɠssŘ ǼÌs ŎŸǣǼ sŘNjÞOÌs_ Ř_ ŎŸǣǼ _sƼĶsǼs_ δˠˤŗɚĶȖsǣʰsǣǼÞŎǼsǣǼÌsǼNjŸƼÌÞOĶsضǼÌʱ
17
ii) δˠˢNNJضsʹNNJʺʰ_Þ¯¯sNjsŘOsEsǼɠssŘǼÌsŎŸǣǼsŘNjÞOÌs_Ř_ǼÌsŎŸǣǼ_sƼĶsǼs_ δˠˢNɚĶȖsǣʰÞǣŎsǣȖNjsŸ¯ǼÌs_ÞɚsNjǣÞǼɴŸ¯EǣĶNjsǣŸȖNjOsǣȖǣs_ʱ
iii) ŎsŘ^ÞǣǼŘOsǼŸNsŘǼNjŸÞ_ʹN^ʺʰɚsNj¶srȖOĶÞ_sŘ_ÞǣǼŘOsŸ¯sOÌǣƼsOÞsǣǼŸ ǼÌs OsŘǼNjŸÞ_ δˠˢN˚δˠˤŗʰ LJȖŘǼÞ¯Þsǣ ǼÌs ǼNjŸƼÌÞO _ÞɚsNjǣÞǼɴ Ř_ ǣƼsOÞsǣ ǣƼOÞض
ɠÞǼÌÞŘǼÌsÞǣŸǼŸƼÞOǣƼOsʱ
iv) ŎsŘ ŗsNjsǣǼ ŗsÞ¶ÌEŸȖNj ^ÞǣǼŘOs ʹŗŗ^ʺʰ rȖOĶÞ_sŘ _ÞǣǼŘOs Ÿ¯ sOÌ ÞŘ_ÞɚÞ_ȖĶ ǼŸ ǼÌs ŘsNjsǣǼ ŘsÞ¶ÌEŸȖNjʰ ƼNjŸɚÞ_sǣ ÞŘ¯ŸNjŎǼÞŸŘ ŸŘ ÞŘ_ÞɚÞ_ȖĶǣ _sŘǣÞǼɴŘ_ƼOĨÞضɠÞǼÌÞŘǼÌsƼŸƼȖĶǼÞŸŘʹǼNjŸƼÌÞONjs_ȖŘ_ŘOɴʺʱ v) ǢǼŘ_Nj_ ^sɚÞǼÞŸŘ Ÿ¯ ǼÌs ŗsNjsǣǼ ŗsÞ¶ÌEŸȖNj ^ÞǣǼŘOs ʹǢ^ŗŗ^ʺ ƼNjŸɚÞ_sǣ ÞŘ¯ŸNjŎǼÞŸŘŸŘǼÌssɚsŘŘsǣǣŸ¯ÞŘ_ÞɚÞ_ȖĶǣƼOĨÞضʳ __ÞǼÞŸŘĶĶɴʰǼŸÞŘɚsǣǼÞ¶ǼsǼÌsǼNjŸƼÌÞOƼǼÌɠɴŸ¯ǼÌsEǣĶǣŸȖNjOsǣǼÌǼǣȖǣǼÞŘʳ ¯ǣOÞǼȖǣ ÞŘ ǼÌs _Þ¯¯sNjsŘǼ ÌEÞǼǼǣʰ DɴsǣÞŘ ŎÞɮÞض ŎŸ_sĶǣ ɠsNjs ƼƼĶÞs_ ǼŸ sǣǼÞŎǼs ǼÌs OŸŘǼNjÞEȖǼÞŸŘ Ÿ¯ sOÌ ǣŸȖNjOsʰ ȖǣÞض ǼÌs NJ ǢÝNJ ƼOĨ¶s ƼNjsɚÞŸȖǣĶɴ ŎsŘǼÞŸŘs_ʳ ǣ EǣĶ ǣŸȖNjOsǣ ɠs OŸŘǣÞ_sNjs_ ĶĶ Ÿ¯ ǼÌs Ÿ¯ ŸNj¶ŘÞO ŎǼǼsNj ǣŸȖNjOsǣ ɚÞĶEĶsÞŘǼÌsǣƼsOÞ¯ÞOÌEÞǼǼʰǼÌǼÞŘOĶȖ_s_ǣs_ÞŎsŘǼNjɴŘ_ƼNjǼÞOȖĶǼsŸNj¶ŘÞO ŎǼǼsNjʹǢŷōŘ_ƻŷōʺʰǣs¶NjǣǣsǣʰͶsʰƼŸŸĶs_ÌĶŸƼÌɴǼsǣŘ_ĶǣŸsƼÞƼÌɴǼsǣ ŸNj ƼĶŘĨǼŸŘ ɠÌsŘ ƼNjsǣsŘǼʳ ǣ ¯ÞǣÌ Njs OŸŘǣÞ_sNjs_ ǣsOŸŘ_ ĶsɚsĶ OŸŘǣȖŎsNjǣʰ ǼÌs ǼNjŸƼÌÞOsŘNjÞOÌŎsŘǼ¯OǼŸNjǣʹǻr®ǣʺȖǣs_¯ŸNjǼÌsŎŸ_sĶɠsNjsŎȖĶǼÞƼĶÞs_EɴǼɠŸŘ_ʰ OOŸNj_ÞضǼŸĶÞǼsNjǼȖNjsɠsNjsʲˡʳˤ̝̈ˠ¯ŸNjδˠˤŗʹəŘ_sNjʉŘ_sŘ˓NJǣŎȖǣǣsŘˡ˟˟ˠʺ
Ř_˟ʳˣ̝̈ˠʳˢ¯ŸNjδˠˢNʹƻŸǣǼˡ˟˟ˡʺʳ
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1.3. NJsǣȖĶǼǣ
ËEÞǼǼǣŎONjŸƼÌɴǼsǣOŸŎƼŸǣÞǼÞŸŘ
^Þ¯¯sNjsŘǼŎONjŸƼÌɴǼsǣOŸŎƼŸǣÞǼÞŸŘǣɠsNjsNjsOŸNj_s_ÞŘǼÌs¯ÞɚsÌEÞǼǼǣsɮŎÞŘs_ Ř_ ǼŸǼĶ Ÿ¯ ˠˣ ǣƼsOÞsǣ ɠsNjs Þ_sŘǼÞ¯Þs_ ʹǻEʳ ˠʺʳ ōÞŘ _Þ¯¯sNjsŘOsǣ ŸOOȖNjNjs_ EsǼɠssŘ ǣǼȖ_ɴ Njsǣʰ ɠÞǼÌÞŘ ɠÌÞOÌEŸǼÌ ǼÌs NjǼÞ¯ÞOÞĶ ÌEÞǼǼǣ ǣÌŸɠs_ ǼÌs ĶŸɠsǣǼ _ÞɚsNjǣÞǼɴ ÞŘ ǼsNjŎǣ Ÿ¯ ŘȖŎEsNj Ÿ¯ ŎONjŸƼÌɴǼsǣ NjsOŸNj_s_ ʹNjsǣƼsOǼÞɚsĶɴ ˡ ÞŘ ǼÌs sɚƼŸNjǼÞŸŘƼŸŘ_ʰrƻÞŘōNjǣĶŘ_ˢÞŘǼÌsNjǼÞ¯ÞOÞĶ_ÞǼOÌʱ^ÞŘəsŘÞOsʺʳƼNjǼ ¯NjŸŎ ǼÌs _sŘǣs ǣs¶Njǣǣ Ŏs_Ÿɠ ʹǢµʺʰ ĶĶ ǼÌs ŸǼÌsNj ÌEÞǼǼǣ ÞŘɚsǣǼÞ¶Ǽs_ ɠsNjs OÌNjOǼsNjÞǣs_ Eɴ ƼǼOÌɴ _ÞǣǼNjÞEȖǼÞŸŘ Ÿ¯ ǼÌs ŎONjŸƼÌɴǼsǣ NjsOŸNj_s_ʰ ɠÞǼÌ ŎONjŸĶ¶s ŸOOȖNjNjÞض ŎÞŘĶɴ ǣ _NjÞ¯Ǽʳ DŸǼÌ ǼÌs ǼɠŸ ÌEÞǼǼǣ Ÿ¯ ǼÌs ǢǼ¶ŘŸŘs _Þ ōNjǣĶ ɠsNjs OÌNjOǼsNjÞǣs_ Eɴ ǼÌs ƼNjsǣsŘOs Ÿ¯ ǼÌs ǣs¶NjǣǣNɴŎŸ_ŸOs ŘŸ_Ÿǣ ǣǣŸOÞǼs_ɠÞǼÌǼÌs¶NjssŘͶNÌsǼŸŎŸNjƼÌĶÞŘȖŎʹŷʳ®ʳōȤĶĶsNjʺħȤǼʊÞضˠ˧ˣˤʳ
ǻEĶsˠʳĵÞǣǼŸ¯ǼÌsǣƼsOÞsǣNjsOŸNj_s_ÞŘǼÌsÌEÞǼǼǣsɮŎÞŘs_ɠÞǼÌÞŘ_ÞOǼÞŸŘŸ¯ǼÌsÞNj ƼNjsǣsŘOsʹ̛ʺŸNjEǣsŘOsʹ˚ʺʳ
Stagnone di Marsala Venice Lagoon
Species Code SG seagrass meadow EP evaporation pond AD artificial ditch SM1 salt marsh SM2 salt marsh Seagrasses
Cymodocea nodosa Cno + + - - -
Ruppia maritima Rma - - + - -
Zostera marina Zma - - - - +
Zostera noltei Zno - - - + +
Macroalgae
Agardhiella subulata Asu - - - - +
Caulerpa prolifera Cpr + - - - -
Chaetomorpha linum Cli + + + - -
Cladophora sp. Cla - - + - -
Cystoseira barbata Cba - - - - +
Glacilaria bursa pastoris Gbp - - - + -
Lomentaria clavellosa Lcl - - - + -
Ulva rigida Uri - - - + +
Halophytes
Atriplex portulacoides - - + - +
Limonium narbonense - - - + -
Sarcocornia fruticosa + + - + +
19
ŎŸŘ¶ ǼÌs ǣs¶Njǣǣsǣ ÞŘ ǼÌs ͶŸŸŘ Ÿ¯ əsŘÞOsʰNJȖƼƼÞ ŎNjÞǼŎ ĵÞŘŘsȖǣ ˠ˦ˤˢ ŸŘĶɴ ŸOOȖNjNjs_ ÞŘ ǼÌs NjǼÞ¯ÞOÞĶ _ÞǼOÌ Ÿ¯ əÞ¶ŘŸĶs ʹ^ʺʰ ɠÌÞĶs ʉŸǣǼsNj ŘŸĶǼsÞ ËŸNjŘsŎŘŘˠ˧ˢˡŸOOȖNjNjs_ÞŘEŸǼÌǼÌsǣĶǼŎNjǣÌÌEÞǼǼǣʹǢōˠŘ_ǢōˡʺǣɠsĶĶ ǣǼÌs¶NjssŘͶȕĶɚNjÞ¶Þ_¶Nj_Ìˠ˧ˡˢʳŎŸŘ¶ǼÌsÌĶŸƼÌɴǼsǣǣƼsOÞsǣƼNjsǣsŘǼ ÞŘ ǼÌs ǣǼȖ_ɴ ǣÞǼsǣʰ ŸŘĶɴ ǼÌs ŎŸǣǼ EȖŘ_ŘǼ ɠsNjs OŸĶĶsOǼs_ Ř_ OŸŘǣÞ_sNjs_ ¯ŸNj ǼNjŸƼÌÞO ǣǣsǣǣŎsŘǼǣʳ ƻNjǼÞOȖĶNjĶɴʰǢNjOŸOŸNjŘÞ ¯NjȖǼÞOŸǣ ʹĵʳʺ ʳ ğʳ ǢOŸǼǼʳ ɠǣ ¯ŸȖŘ_ ɠÞ_sǣƼNjs_ÞŘĶŎŸǣǼĶĶŸ¯ǼÌsÌEÞǼǼǣÞŘɚsǣǼÞ¶Ǽs_ʰOŸɚsNjÞضŎŸǣǼŸ¯ǼÌsEŸNj_sNjǣ Ÿ¯ǼÌsǣĶǼŎNjǣÌsǣʳ ®ÞǣÌ_sŎŸ¶NjƼÌÞOǼNjÞǼǣ ®ÞǣÌǣǣsŎEͶsǣ˚^Þ¯¯sNjsŘǼ¯ÞǣÌǣǣsŎEͶsǣŸOOȖNjNjs_ŎŸŘ¶ĶĶŸ¯ǼÌsÌEÞǼǼǣ ÞŘɚsǣǼÞ¶Ǽs_ʰɠÞǼÌǣŸŎsɚNjÞEÞĶÞǼɴǼǣsǣŸŘĶĶsɚsĶʹǻEʳˡʺʳŷɚsNjĶĶʰǼŸǼĶŸ¯ˠˡ ǣƼsOÞsǣEsĶŸŘ¶ÞضǼŸ˥¯ŎÞĶÞsǣɠsNjsǣŎƼĶs_ʰŎŸǣǼŸ¯ɠÌÞO̯sĶĶÞŘǼÌssOŸĶŸ¶ÞOĶ ¶NjŸȖƼŸ¯sǣǼȖNjÞŘsNjsǣÞ_sŘǼŘ_ÞŘɚsNjǼÞɚŸNjŸȖǣŸNjŸŎŘÞɚŸNjŸȖǣǣ¯ss_Þض¶NjŸȖƼʳǻÌs ÌÞ¶ÌsǣǼŘȖŎEsNjŸ¯ǣƼsOÞsǣɠǣ¯ŸȖŘ_ÞŘǼÌsÞŘŘsNjǣĶǼŎNjǣÌʹǢōˡʺʳ ǻÌsǼNj¶sǼǣƼsOÞsǣʳ¯ǣOÞǼȖǣǣÌŸɠs_OŸŎŎŸŘǼNjsŘ_ÞŘŎŸǣǼŸ¯ǼÌsÌEÞǼǼǣʲÞǼ ɠǣ_ŸŎÞŘŘǼÞŘǣƼNjÞضʰEŸǼÌÞŘǼsNjŎǣŸ¯_sŘǣÞǼɴʹŘȖŎEsNjŸ¯ÞŘ_ÞɚÞ_ȖĶǣŎ˚ˡʺŘ_ ƼsNjOsŘǼ¶sEȖŘ_ŘOsʹ̇ʺŘ__sONjsǣs_ÞŘȖǼȖŎŘʳ rɮOsƼǼÞŸŘǣɠsNjs¶ÞɚsŘEɴǼÌsNjǼÞ¯ÞOÞĶOÌŘŘsĶʹ^ʺŘ_ǼÌsÞŘŘsNjǣĶǼŎNjǣÌʹǢōˡʺʰ ɠÌsNjsʳ¯ǣOÞǼȖǣ_sŘǣÞǼÞsǣɠsNjsÌÞ¶ÌsNjsɚsŘÞŘȖǼȖŎŘŘ_EɴǼÌs¯ÞNjǣǼǣĶǼŎNjǣÌ ǣÞǼsʹǢōˠʺʰɠÌŸǣs OŸŎŎȖŘÞǼɴɠǣ_ŸŎÞŘǼs_EɴǼÌsNjÞŘEŸɴsNjÞʹNJÞǣǣŸʰˠ˧ˠ˟ʺÞŘ ǣƼNjÞضŘ_EɴĵÞʊǣĶÞsŘǣʹNJÞǣǣŸʰˠ˧ˠ˟ʺÞŘȖǼȖŎŘʳʳEŸɴsNjÞɠǣĶǣŸ_ŸŎÞŘŘǼÞŘ ȖǼȖŎŘɠÞǼÌÞŘǼÌs ǣÞǼsǣÞŘǼÌsǢǼ¶ŘŸŘs_ÞōNjǣĶʹǢµŘ_rƻʺŘ_ǢōˡÞŘǼÌs əsŘÞOsͶŸŸŘʳ
ʳ ¯ǣOÞǼȖǣ_sŎŸ¶NjƼÌÞO ǣǼNjȖOǼȖNjs ˚ǻÌs _sŎŸ¶NjƼÌÞO ǣǼNjȖOǼȖNjs Ÿ¯ ǼÌs ǼNj¶sǼ ƼŸƼȖĶǼÞŸŘǣ ɠǣ ÞŘɚsǣǼÞ¶Ǽs_ ǼÌNjŸȖ¶Ì ¯NjsLJȖsŘOɴ _ÞǣǼNjÞEȖǼÞŸŘǣ Ÿ¯ ǼÌs ǣǼŘ_Nj_ ĶsضǼÌʹ®Þ¶ʳˡʺʳĶĶǼÌsƼŸƼȖĶǼÞŸŘǣǣÌŸɠs__ÞǣǼNjÞEȖǼÞŸŘǣǣĨsɠs_ǼŸǼÌsĶŸɠsNjǣÞʊs OĶǣǣsǣʹŎÞŘĶɴǢĵ̣ˣ˟ŎŎʺʳǻÌsŸŘĶɴsɮOsƼǼÞŸŘɠǣ¶ÞɚsŘÞŘǢōˠÌEÞǼǼʰɠÌsNjs
20 ¯sɠ ÞŘ_ÞɚÞ_ȖĶǣ ɠsNjs ȖŘÞ¯ŸNjŎĶɴ _ÞǣǼNjÞEȖǼs_ʳ ǻÌs ǣsǣŸŘĶ ǼNjsŘ_ ǣssŎǣ ǼŸ ¯ŸĶĶŸɠ ŸƼƼŸǣÞǼs ƼǼǼsNjŘǣ ɠÞǼÌÞŘ ǼÌs ǼɠŸ ǣǼȖ_ɴ Njsǣʲ _sǣƼÞǼs ǼÌs ŘsǼ Njs_ȖOǼÞŸŘ Ÿ¯ ÞŘ_ÞɚÞ_ȖĶǣ¯NjŸŎǣƼNjÞضǼŸȖǼȖŎŘʰÞǼÞǣOĶsNjÌŸɠǼÌsŎŸ_sǣÌÞ¯Ǽǣ¯NjŸŎĶŸɠsNjǼŸ ǣĶÞ¶ÌǼĶɴÌÞ¶ÌsNjOĶǣǣsǣÞŘōNjǣĶʹǢµŘ_rƻʺʰŘ_ɚÞOsɚsNjǣÞŘəsŘÞOsʹǢōˠʰǢōˡ Ř_^ʺʳ
®Þ¶ȖNjs ˡʳ ®NjsLJȖsŘOɴ _ÞǣǼNjÞEȖǼÞŸŘǣ Ÿ¯ ǼÌs ǣǼŘ_Nj_ ĶsضǼÌ Ÿ¯ʳ ¯ǣOÞǼȖǣ ƼŸƼȖĶǼÞŸŘǣ OŸĶĶsOǼs_ÞŘǼÌsǣŎƼĶÞضǣÞǼsǣÞŘǣƼNjÞضʹɠÌÞǼsENjǣʺŘ_ȖǼȖŎŘʹŸɚsNjĶƼƼs_¶NjsɴENjǣʺʳ
ǻÌs ǣsɮ NjǼÞŸ EsǼɠssŘ ¯sŎĶsǣ Ř_ ŎĶsǣ NjsOŸNj_s_ ɠǣ ÌÞ¶ÌĶɴ ɚNjÞEĶs ʹǻEʳ ˢʺ ONjŸǣǣ ÌEÞǼǼǣ Ř_ ǣsǣŸŘǣʰ ɠÞǼÌ ¶sŘsNjĶ ǼsŘ_sŘOɴ ǼŸ ¯sŎĶs ƼNjsɚĶsŘOsʰ ɠÞǼÌ ǼÌssɮOsƼǼÞŸŘŸ¯ǼÌsNjǼÞ¯ÞOÞĶ_ÞǼOÌÞŘəÞ¶ŘŸĶsʰɠÌsNjsŎĶsǣɠsNjsŎŸNjsEȖŘ_ŘǼÞŘ EŸǼÌǣŎƼĶÞضǣsǣŸŘǣʳ
ǻEĶsˢʳǢsɮNjǼÞŸ®sŎĶsʲōĶsŸ¯ʳ¯ǣOÞǼȖǣOŸĶĶsOǼs_ ÞŘEŸǼÌǣŎƼĶÞضǣsǣŸŘǣONjŸǣǣÌEÞǼǼǣʳ SG EP AD SM1 SM2 Spring 16.08 0.90 0.53 9.50 1.25 Autumn 3.33 211.00 0.52 3.78 24.20 Stagnone di Marsala F:M Venice lagoon
21 ǻEĶs ˡʳ ^ sŘǣÞǼɴ ʹŘȖŎE sNj Ÿ¯ ÞŘ_Þ ɚÞ_ȖĶǣ ˠ˟ ˟ Ŏ ˚ˡ ʺ Ř_ ƼsNjO sŘǼ¶s EȖŘ_ ŘOs ʹ̇ʺ Ÿ¯ ǼÌs ǣƼsO Þsǣ ǣŎƼĶs_ ÞŘ ǼÌs ǣǼ Ȗ_ɴ ǣÞǼsǣ ÞŘ ǣƼNjÞض Ř_ ȖǼȖŎŘʳ rOŸĶŸ¶ÞO Ķ Ř_ ¯ss_Þض ¶NjŸȖƼǣ Njs ĶǣŸ ÞŘ_Þ OǼs_ OOŸNj_ Þض ǼŸ ®Nj ŘOŸ sǼ Ķʳ ʹˡ˟ ˟˥ ʺ Ř_ ĵs ŸŘNj_Ÿǣ ʹˡ˟ ˟˧ ʺǣƼsO Þ¯ÞO ĶĶɴ ¯ŸNj ʳ ¯ ǣOÞǼȖǣ ʳ rOŸĶŸ¶ÞO Ķ ¶NjŸȖƼǣ Njs ʲrNJ ʰs ǣǼȖ NjÞŘs NjsǣÞ_sŘǼʰōğ ʰŎNjÞŘs ĠȖɚsŘ ÞĶs ŎÞ ¶Nj ŘǼʰōǢʰ Ŏ Nj ÞŘsǣsǣŸŘ Ķ Ŏ Þ¶ NjŘǼʳ ǻNjŸ ƼÌÞ O ¶NjŸȖƼǣ Njsʲ N Ǣʰ ONjŘÞɚ ŸNjsǣʰ ÝǢʰ ÞŘɚsNjǼÞɚŸNjsǣʰÝ ®ʰÞŘɚsNjǼsENj Ǽs˚¯Þǣ̯ss_s NjǣʰË Nʰ ÌsNj EÞɚŸNjsǣ˚ONjŘÞɚŸNj sǣʰŷʰŸ Ŏ ŘÞɚŸNjsǣʳ Fa m ily Sp e cie s Ec ol ogi ca l gr oup Feed in g gr oup Se as o n De n si ty % De n si ty % De n si ty % De n si ty % De n si ty % Sp ri n g 10. 0 3 .8 58. 7 26. 5 -134. 4 83. 1 -Au tu m n 136. 3 73. 6 190. 0 82. 6 -2 .0 52. 2 Sp ri n g 256. 3 96. 2 144. 6 65. 2 231. 9 65. 3 27. 3 16. 9 23. 4 27. 3 Au tu m n 48. 8 26. 4 27. 5 12. 0 58. 8 50. 5 15. 7 31. 0 157. 5 90. 0 Sp ri n g - -Au tu m n -0. 7 17. 4 Sp ri n g - -Au tu m n - ----0 .2 4 .3 Sp ri n g -123. 1 34. 7 -Au tu m n -57. 5 49. 5 -5 .0 2. 9 Sp ri n g -8. 8 10. 2 Au tu m n - ----0 .3 8 .7 Sp ri n g -14. 0 16. 3 Au tu m n - -Sp ri n g - ----3 .2 3 .8 Au tu m n -34. 7 68. 5 0 .5 13. 0 Sp ri n g - -Au tu m n -0 .3 0 .6 -Sp ri n g - ----0 .6 0 .8 Au tu m n - -Sp ri n g -9 .8 4 .4 -Au tu m n -3 .8 1 .6 -Sp ri n g -7 .6 3 .4 -Au tu m n -8 .8 3 .8 ----0 .2 4 .3 Syngna thi d a e Sy ngnat h us abas te r ER IS Syngna thi d a e Sy ngnat h us t y p hl e ER IF Mu g ilid ae M u gi l c e phal us MJ O Spa ri d a e Spar us a ur at a MJ O Mu g ilid ae Li za r am ada MJ O Mu g ilid ae Li za s a lien s ER H C G o b iid ae K n ip ow it sc hi a pani zz a e ER IS Mu g ilid ae Li za a u ra ta MS H C G o b iid ae P o m a to sc hi st us m ar m o ra tu s ER IS G o b iid ae P o m a to sc hi st us m in ut us MJ IS A the ri ni da e A the ri na b oy e ri ER C S C y pri n odonti da e A p hani us f as ci at us ER O St ag n o n e d i M ar sa la V e n ic e la go o n SG Se ag ra ss m e ad o w EP Ev ap or at io n pond AD A rt if ici al d it ch SM 1 Sa lt m ar sh SM 2 Sa lt m ar sh
22 ÝǣŸǼŸƼÞONjsǣȖĶǼǣ
ŷNj¶ŘÞOŎǼǼsNjǣŸȖNjOsǣŘ_¯ÞǣÌOŸŎŎȖŘÞǼÞsǣ
ǻÌs ŸɚsNjĶĶ Njضsǣ Ÿ¯ EŸǼÌ δˠˢN Ř_ δˠˤŗ ǼÌǼ ÞŘOĶȖ_s ĶĶ ǼÌs LJȖǼÞO ǣŸȖNjOsǣ Ÿ¯
ŸNj¶ŘÞOŎǼǼsNjɚÞĶEĶsÞŘǼÌsÌEÞǼǼǣǣŎƼĶs_ʹŘŎsĶɴsɮOĶȖ_ÞضǼÌsǼsNjNjsǣǼNjÞĶ ÌĶŸƼÌɴǼsǣʺ ǣÌŸɠs_ ɚĶȖsǣ ŎŸNjs _sƼĶsǼs_ ÞŘ ǼÌs əsŘÞOs ĵ¶ŸŸŘ ǼÌŘ ÞŘ ǼÌs ǢǼ¶ŘŸŘs_ÞōNjǣĶʹ®Þ¶ʳˢʺʳǣÞ¶ŘÞ¯ÞOŘǼǣÌÞ¯ǼɠǣNjsOŸNj_s_¯ŸNjδˠˢNEsǼɠssŘǼÌs ǼɠŸǣǼȖ_ɴNjsǣʹǼ˚ǼsǣǼʰƼ˚ɚĶȖṣˡʳˡs˚ˠ˥ʺŘ_ɠÞǼÌÞŘsOÌͶŸŸŘʰEsǼɠssŘŘǼȖNjĶ Ř_NjǼÞ¯ÞOÞĶÌEÞǼǼǣʹħNjȖǣĨĶ˚ɟĶĶÞǣƼŸǣǼ˚ÌŸOǼsǣǼʰƼ˚ɚĶȖṣ˟ʳ˟ˤʺʳÝŘǼÌsǢǼ¶ŘŸŘsʰ ǼÌs ŎsŘ Ÿ¯ ONjEŸŘ ǣÞ¶ŘǼȖNjsǣ Njضs_ ¯NjŸŎ ˚ˠ˦ʳ˧ˠ̝˟ʳˣˠ ǼŸ ˚˦ʳˣˤ̝˟ʳˣ˦̈ ÞŘ ǼÌs ǣs¶Njǣǣ Ŏs_Ÿɠ ʹǢµʺ Ř_ ¯NjŸŎ ˚ˡ˟ʳ˦˦̝ˠʳ˟ˠ ǼŸ ˚˨ʳˠ˧̝˟ʳˢ̈ ÞŘ ǼÌs ǣĶǼ sɚƼŸNjǼÞŸŘ ƼŸŘ_ ʹrƻʺʳ ÝŘ ǼÌs əsŘÞOs ĵ¶ŸŸŘʰ ÞŘǣǼs_ʰ ǼÌs Njضsǣ ɚNjÞs_ ¯NjŸŎ NjŸȖŘ_˚ˡˢǼŸEŸȖǼ˚ˠ˟ʳˤ̈ÞŘEŸǼÌǣĶǼŎNjǣÌÌEÞǼǼǣʹǢōˠŘ_OǢōˡʺŘ_ɠsNjs sɚsŘ ŎŸNjs _sƼĶsǼs_ ÞŘ ǼÌs NjǼÞ¯ÞOÞĶ _ÞǼOÌ Ÿ¯ əÞ¶ŘŸĶs ʹ^ʺʰ NjضÞض ¯NjŸŎ ˚ ˢ˟ʳˢˠ̝ˠʳˣ˨ǼŸ˚ˠ˦ʳˠ˥̝˟ʳˣ˧̈ʳ EŸȖǼǼÌsδˠˤŗʰŘŸsɚÞ_sŘǼ_Þ¯¯sNjsŘOsǣsŎsNj¶s_EsǼɠssŘͶŸŸŘǣʰEȖǼ^ÞŘǼÌs
əsŘÞOs ĵ¶ŸŸŘ ɠǣ ǣÞ¶ŘÞ¯ÞOŘǼĶɴ ǣÌÞ¯Ǽs_ ʹħNjȖǣĨĶ˚ɟĶĶÞǣ ƼŸǣǼ˚ÌŸO ǼsǣǼʰ Ƽ˚ ɚĶȖṣ˟ʳ˟ˤʺǼŸɠNj_ǣĶŸɠsNjNjضsʹ¯NjŸŎˢʳˠ̝˟ʳˠ˧ǼŸ˨ʳ˨̝˟ʳˡˤ̈ʺǼÌŘÞŘ ĶĶǼÌs ŸǼÌsNjÌEÞǼǼǣʰǼÌǼNjضs_ƼƼNjŸɮÞŎǼsĶɴ¯NjŸŎˣǼŸˠˡ̈ʳ
ËĶŸƼÌɴǼsǣǣÞ¶ŘǼȖNjsǣĶɠɴǣ¯sĶĶɠÞǼÌÞŘǼÌsŗNjضsʰEȖǼɠsNjsŎŸNjs_sƼĶsǼs_ÞŘNʰ ɠÞǼÌǼÌssɮOsƼǼÞŸŘŸ¯^ʰɠÌsNjsÞǼɠǣƼĶOs_EsǼɠssŘǼÌsǣÞ¶ŘǼȖNjsǣŸ¯ƻŷōŘ_ Ǣŷōʳ ǻÌs ǣÌÞ¯Ǽ ŸEǣsNjɚs_ ¯ŸNj ǼÌs LJȖǼÞO ǣŸȖNjOsǣ Ÿ¯ ŸNj¶ŘÞO ŎǼǼsNj ɠǣ ĶǣŸ ŎÞŘǼÞŘs_ Ǽ ¯ÞǣÌ OŸŎŎȖŘÞǼɴ ĶsɚsĶʰ ɠÌŸǣs ŸɚsNjĶĶ Njضsǣʰ sǣƼsOÞĶĶɴ Ÿ¯ ONjEŸŘ ÞǣŸǼŸƼÞOǣÞ¶ŘǼȖNjsǣ¯sĶĶɠÞǼÌÞŘŎŸNjs_sƼĶsǼs_ɚĶȖsǣÞŘəsŘÞOsĵ¶ŸŸŘŘ_ɠÞǼÌÞŘ sOÌͶŸŸŘʰǼÌsɴɠsNjsǣǼÞĶĶŎŸNjs_sƼĶsǼs_ÞŘǼÌsNjǼÞ¯ÞOÞĶÌEÞǼǼǣʹ®Þ¶ʳˢʺʳ
23 ®Þ¶ȖNjsˢʳδˠˢNʹ̈ʺɚǣʳδˠˤŗʹ̈ʺŸ¯¯ÞǣÌǣƼsOÞsǣŘ_ŘŘȖĶŎsŘʹ̝ǣʳ_ʳʺŸ¯ŸNj¶ŘÞOŎǼǼsNj ǣŸȖNjOsǣǣŎƼĶs_ÞŘsOÌÌEÞǼǼǣÞǼsʲǢµʰǣs¶NjǣǣŎs_ŸɠŘ_rƻʰsɚƼŸNjǼÞŸŘƼŸŘ_ÞŘ ǢǼ¶ŘŸŘs_ÞōNjǣĶʱǢōˠŘ_ǢōˡʰǣĶǼŎNjǣÌsǣʰ^ʰNjǼÞ¯ÞOÞĶ_ÞǼOÌÞŘəsŘÞOsĵ¶ŸŸŘʳ®ȖĶĶ OŸĶŸȖNjs_ƼŸÞŘǼǣNjsǣÞضĶsÞŘ_ÞɚÞ_ȖĶǣŸ¯¯ÞǣÌǣƼsOÞsǣʰEĶOĨǣɴŎEŸĶǣNjsŸNj¶ŘÞOŎǼǼsNj ǣŸȖNjOsǣʰOOŸNj_ÞضǼŸǼÌsĶs¶sŘ_ǣƼsOÞ¯Þs_ÞŘǼÌs¶NjƼÌʳ
24
NŸŎƼNjÞǣŸŘŸ¯ʳ¯ǣOÞǼȖǣÞǣŸǼŸƼÞOŘÞOÌsɠÞǼÌÞŘÌEÞǼǼǣ
ǻÌs ÞǣŸǼŸƼÞO ŘÞOÌs Ÿ¯ʳ ¯ǣOÞǼȖǣ ɚNjÞs_ ÞŘ ɠÞ_ǼÌʰ ǣÌƼs Ř_ ƼŸǣÞǼÞŸŘ ONjŸǣǣ ÌEÞǼǼǣŘ_ǣsǣŸŘǣʹ®Þ¶ʳˣʺǣɠsĶĶǣÞǼǣ_sǣONjÞƼǼÞɚsŎsǼNjÞOǣʹǻEʳˣʺʳ
ǣ ¶sŘsNjĶ ǼNjsŘ_ʰ ÞŘ ĶĶ ǼÌs ÌEÞǼǼǣ ÞŘɚsǣǼÞ¶Ǽs_ʰ ǼÌs ŎsǼNjÞOǣ _sǣONjÞEÞض ǼÌs ÞǣŸǼŸƼÞO ŘÞOÌs Ÿ¯ʳ ¯ǣOÞǼȖǣ ɠsNjs ÌÞ¶ÌsNj ÞŘ ȖǼȖŎŘ ǼÌŘ ÞŘ ǣƼNjÞض ʹǻEʳ ˣʺʰ ÞŘ_ÞOǼÞضÌÞ¶ÌsNjsɮǼsŘǣÞŸŘŸ¯ǼÌsÞǣŸǼŸƼÞOŘÞOÌsʹǢrOʰŗNJŘ_NNJʺʰŘ_ĶǣŸ ÌÞ¶ÌsNjǼNjŸƼÌÞO_ÞɚsNjǣÞǼɴŸ¯ǼÌsÞŘ_ÞɚÞ_ȖĶǣʹN^ʺʰNjs_ȖŘ_ŘOɴʹŗŗ^ʺŘ_sɚsŘŘsǣǣŸ¯ ǼÌsÞŘ_ÞɚÞ_ȖĶǣƼOÞضɠÞǼÌÞŘǼÌsŘÞOÌsʹǢ^ŗŗ^ʺʳǻÌsŸŘĶɴsɮOsƼǼÞŸŘɠǣ¶ÞɚsŘÞŘ rƻʰɠÌsNjsŗNJŘ_NNJɠsNjsĶŸɠsNjÞŘȖǼȖŎŘǼÌŘÞŘǣƼNjÞضʳ
ǻEĶs ˣʳ ƻŸƼȖĶǼÞŸŘ˚ɠÞ_s ŎsǼNjÞOǣ OĶOȖĶǼs_ ¯ŸNj sOÌ ǣƼsOÞsǣ Ř_ ÌEÞǼǼ ÞŘ ǣƼNjÞض Ř_ ȖǼȖŎŘʳ SEAc NR CR CD NND SDNND SEAc NR CR CD NND SDNND Ǣµ A. fasciatus 1.04 2.44 1.24 0.74 0.24 0.11 1.91 3.89 1.64 1.28 0.47 0.31 A. boyeri 0.55 1.22 0.86 0.42 0.39 0.34 0.10 0.43 0.42 0.20 0.15 0.09 rƻ A. fasciatus 1.15 2.82 2.01 0.70 0.32 0.37 1.54 2.26 1.43 0.79 0.51 0.43 A. boyeri 0.83 3.11 1.20 0.67 0.43 0.42 1.10 2.55 1.68 0.73 0.49 0.38 S. abaster 1.10 1.48 1.17 0.56 0.56 0.35 4.25 1.66 1.68 0.98 1.49 0.20 S.typhle 1.10 1.92 1.18 0.71 0.59 0.54 2.21 1.90 2.14 1.20 1.18 1.21 ^ A. fasciatus 3.28 2.75 4.72 1.38 0.39 0.23 6.00 3.59 7.28 2.24 0.51 0.25 K. Panizzae 1.43 1.39 1.97 0.81 0.56 0.12 0.57 0.67 3.02 0.65 0.36 0.29 Ǣōˠ A. fasciatus 1.00 1.95 2.40 0.64 0.39 0.32 4.58 3.43 5.52 1.78 0.89 0.50 A. boyeri 3.94 2.57 3.40 1.41 0.42 0.60 - - - -L. saliens - - - 0.59 0.81 1.68 0.50 0.43 0.27 Ǣōˡ A. fasciatus 1.37 1.64 2.62 0.87 0.42 0.17 2.91 3.96 3.56 1.35 0.70 0.40 A. boyeri - - - 4.56 3.14 5.03 1.58 1.14 0.68 K. Panizzae - - - 0.16 1.48 0.20 0.37 0.34 0.24 L. aurata 2.49 1.48 2.97 1.14 0.68 0.33 - - - -L. saliens 1.34 1.57 2.43 0.98 0.47 0.19 1.97 2.71 2.78 1.41 0.79 0.16 P. marmoratus - - - 0.61 1.17 1.48 0.51 0.35 0.20 Spring Autumn
25 OŸŎƼNjsÌsŘǣÞɚsĶŸŸĨǼǼÌsŎsǼNjÞOǣŸ¯ǼÌsɠÌŸĶsOŸŎŎȖŘÞǼÞsǣɠÌsNjsʳ¯ǣOÞǼȖǣ ɠǣƼĶOs_ʹǻEʳˤʺǣÌŸɠs_ǣÞŎÞĶNjǼNjsŘ_ɠÞǼÌÞŘEŸǼÌͶŸŸŘǣʰɠÞǼÌǼÌsNjǼÞ¯ÞOÞĶ ÌEÞǼǼǣʰrƻŘ_^ƼNjsǣsŘǼÞضǼÌsɠÞ_sǣǼÞǣŸǼŸƼÞOŘÞOÌsʹÌÞ¶ÌsNjǢrOɚĶȖsǣʺŘ_ ǼÌsÌÞ¶ÌsǣǼǼNjŸƼÌÞO_ÞɚsNjǣÞǼɴʹÌÞ¶ÌsNjN^ʺʳ
ǻEĶsˤʳNŸŎŎȖŘÞǼɴ˚ɠÞ_sŎsǼNjÞOǣŸ¯ɠÌŸĶsOŸŎŎȖŘÞǼÞsǣǣŎƼĶs_ǼsOÌÌEÞǼǼǣÞǼsʳ
SEAc NR CR CD NND SDNND SG 1.54 4.81 2.43 0.89 0.24 0.22 EP 3.31 5.39 5.42 1.28 0.27 0.27 AD 7.89 4.98 8.76 2.56 0.37 0.19 SM1 7.13 5.66 6.24 2.19 0.44 0.37 SM2 6.58 4.76 7.82 1.88 0.36 0.28
ÝŘ ŎŸǣǼ Ÿ¯ ǼÌs ÌEÞǼǼǣʰ ǼÌs ÞǣŸǼŸƼÞO ŘÞOÌs Ÿ¯ ǼÌs ǼNj¶sǼ ǣƼsOÞsǣ ɠǣ OĶsNjĶɴ ǣsƼNjǼs_¯NjŸŎǼÌsŘÞOÌsǣŸ¯ǼÌsOŸ˚ŸOOȖNjNjÞضǣƼsOÞsǣŘ_ĶŸOǼs_ÞŘǼŸŎŸNjsˠˢN
Ř_ˠˤŗ˚sŘNjÞOÌs_NjsŸ¯ǼÌsÞǣŸǼŸƼÞOǣƼOsʹ®Þ¶ʳ ˣʺʳOŸŘǣÞ_sNjEĶs ŸɚsNjĶƼɠÞǼÌ
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